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Cyperaceae source details

Larridon, I.; Rabarivola, L.; Xanthos, M.; Muasya, A. M. (2019). Revision of the Afro-Madagascan genus Costularia (Schoeneae, Cyperaceae): infrageneric relationships and species delimitation. PeerJ. 7: e6528.
497239
10.7717/peerj.6528 [view]
Larridon, I.; Rabarivola, L.; Xanthos, M.; Muasya, A. M.
2019
Revision of the Afro-Madagascan genus <i>Costularia</i> (Schoeneae, Cyperaceae): infrageneric relationships and species delimitation
PeerJ
7: e6528
Publication
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2024-10-22 06:12:12Z
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2024-10-23 07:51:15Z
changed

Costularia andringitrensis Larridon (original description)
Costularia cadetii Larridon (original description)
Costularia itremoensis Larridon (original description)
Costularia pantopoda var. pantopoda (Baker) C.B.Clarke ex Cherm. (original description)
Costularia robusta (Cherm.) Larridon (new combination reference)
Holotype TAN, geounit Madagascar, identified as Costularia andringitrensis Larridon
Lectotype K 000244883, geounit Madagascar, identified as Costularia leucocarpa (Ridl.) H.Pfeiff.
Lectotype K 000244884, geounit Madagascar, identified as Cladium fimbristyloides Baker
Lectotype K 000244885, geounit Madagascar, identified as Costularia baronii C.B.Clarke
Lectotype K 000244886, geounit Madagascar, identified as Cladium pantopodum Baker
Lectotype K 000244887, geounit Madagascar, identified as Costularia pilisepla (Steud.) J.Kern
Lectotype K 000244888, geounit Madagascar, identified as Cladium melleri Baker
Lectotype K 000244893, geounit KwaZulu-Natal, identified as Costularia natalensis C.B.Clarke
Holotype K 001418184, geounit Réunion, identified as Costularia cadetii Larridon
Holotype P 00459966, geounit Madagascar, identified as Costularia robusta (Cherm.) Larridon
Lectotype P 00459967, geounit Madagascar, identified as Costularia laxa var. macrantha Cherm.
Lectotype P 00459974, geounit Madagascar, identified as Costularia brevifolia Cherm.
Lectotype P 00459980, geounit Madagascar, identified as Costularia humbertii Bosser
Lectotype P 00459983, geounit Madagascar, identified as Costularia laxa Cherm.
Isolectotype P 00459985, geounit Madagascar, identified as Rhynchospora leucocarpa Ridl.
Isolectotype P 00459986, geounit Madagascar, identified as Cladium fimbristyloides Baker
Isolectotype P 00459987, geounit Madagascar, identified as Cladium melleri Baker
Isolectotype P 00459988, geounit Madagascar, identified as Cladium pantopodum Baker
Isolectotype P 00459989, geounit Madagascar, identified as Costularia baronii C.B.Clarke
Lectotype P 00552880, geounit Mauritius, identified as Cyperus melicoides Poir.
Lectotype P 00723561, geounit Madagascar, identified as Costularia baronii var. microcarpa Cherm.
Isotype UPOS 008893, geounit Réunion, identified as Costularia cadetii Larridon
Holotype P 0318446, geounit Madagascar, identified as Costularia itremoensis Larridon
 Description

Perennial herbs, small to tall, tufted or more rarely shortly rhizomatous, caudex sometimes present. Culms scapose ... [details]

 Description

Small perennial herb, flowering culm up to 24 cm, scarcely exceeding the leaves. Caudex absent. Culm (excluding the ... [details]

 Description

Robust perennial herb: Culms 20–80 cm × c. 5 mm, generally quite robust. Basal leaves with leaf blades 40–80 ... [details]

 Description

Robust perennial herb: Caudex 10–12 cm × 4–5 cm. Culm (appearing) lateral, robust, 50–80 cm × 5–8 mm, ... [details]

 Description

Small perennial herb, flowering culm up to 28 cm, scarcely exceeding the leaves. Caudex absent or short (c. 0.5 mm ... [details]

 Description

Robust perennial herb: Caudex robust, 5–10 cm × 1–2 cm. Culm 30–70 cm × 2–3 mm, compressed, smooth. Basal ... [details]

 Description

Medium-sized to tall perennial herb, up to c. 1.4 m. Culm 17–68 cm × 1.7–3.5 mm. Basal leaves distichous; leaf ... [details]

 Description

Perennial herb up to c. one m tall with a woody rhizome (c. four mm diam.), caudex sometimes present (c. seven mm ... [details]

 Description

Perennial herb with short rhizome with stiff fibres. Culm 35–100 cm × 1.5–4 mm, striate, minutely ... [details]

 Description

Very robust and tall perennial herb: Culm robust, 1.2–2 m × c. 6 mm. Basal leaves firm, long, five mm wide, ... [details]

 Description

Perennial herb up to 2.5 m tall, tufted; rhizome 1–1.5 mm in diameter, woody, erect, clothed in thick adventitious ... [details]

 Description

Culms slender, 0.9 mm wide. Basal leaves with the leaf sheath little larger than the narrow leaf blades (up to c. ... [details]

 Description

Medium-sized perennial herb, up to c. 65 cm height. Culm 25–30 cm × 2.5–3 mm. Basal leaves distichous; leaf ... [details]

 Description

Perennial herb up to 2.5 m tall. Caudex covered in lateral roots can be present (0.7–1.5 cm in diam.). Culms more ... [details]

 Description

Perennial herb up to 2.5 m tall, caespitose, forming dense clumps. Culm c. 80 cm × 3.5–5 mm, cylindrical, ... [details]

 Distribution

Southeastern Africa (Malawi, Mozambique, South Africa, Swaziland, Zimbabwe), Madagascar, the Mascarenes (La ... [details]

 Distribution

The species is only known from south-central Madagascar, where it was found in the Andringitra National Park, Haute ... [details]

 Distribution

Costularia baronii occurs in the Antananarivo, Fianarantsoa and Toliara provinces of Madagascar. [details]

 Distribution

Endemic to southeastern Madagascar and only known from the Atsimo Atsinana and Anosy regions in the Fianarantsoa ... [details]

 Distribution

Costularia cadetii is a small perennial herb, endemic to La Réunion and found only in the Parc National de La ... [details]

 Distribution

Endemic to the Antsiranana province of Madagascar where it is restricted to the high-elevation zone of the Marojejy ... [details]

 Distribution

Costularia itremoensis is endemic to Madagascar and is found in the highlands of South Central Madagascar, in the ... [details]

 Distribution

Endemic to Madagascar, found in the provinces Antananarivo, Antsiranana, Fianarantsoa, Toamasina and Toliara. [details]

 Distribution

Endemic to the Mascarene Islands of La Réunion and Mauritius. [details]

 Distribution

Endemic to Madagascar, occurring in the provinces Antananarivo, Fianarantsoa and Toamasina. [details]

 Distribution

Endemic to the Ihorombe region of Fianarantsoa province in Madagascar. [details]

 Distribution

Costularia natalensis is restricted in its distribution to southeastern Africa (Fig. 16). In particular, the ... [details]

 Distribution

Costularia pantopoda var. pantopoda occurs in the Fianarantsoa province and in the south of the Antananarivo ... [details]

 Distribution

Costularia purpurea is endemic to Madagascar, occurring in the Antananarivo, Antsiranana, Fianarantsoa, Toamasina ... [details]

 Distribution

Costularia robusta is only known from the Manongarivo, Marojejy and Tsaratanana protected areas and their environs ... [details]

 Distribution

Costularia xipholepis is endemic to the Seychelles and restricted to Mahé and has been found in three locations ... [details]

 Ecology

This species is found in near rocks in grassland to ericoid shrubland vegetation at 2,000–2,500 m in ... [details]

 Ecology

It has been found growing in rocky areas (e.g. rock crevices along a stream bank), ericoid shrubland at elevations ... [details]

 Ecology

It has been found growing on humid rocks in peatlands, on laterite and granite in tropical forest, and in faults of ... [details]

 Ecology

Found growing in rocky areas, montane grasslands and ericoid vegetation close to volcanic crater edges at high ... [details]

 Ecology

Found growing in swamps in high elevation ericoid vegetation, and on gneiss and quartzite rocks of the mountain ... [details]

 Ecology

The habitat in which this species is found consists of bare rocks and/or grassland in the Itremo massif (L. ... [details]

 Ecology

The species occurs at mid to (very) high elevation, and has been collected along mountain ridges, from thickets on ... [details]

 Ecology

Costularia melicoides prefers mid to higher elevation on the island of La Réunion: (500–)900–1,700 (–2,000) ... [details]

 Ecology

Marshes, humid areas in forest, an elevation of 1,000–1,500 m. Specimens with very young inflorescences were ... [details]

 Ecology

Found growing on shaded, humid sandstone in Isalo National Park at an elevation of c. 1,000 m. Only known specimen ... [details]

 Ecology

Ppopulations are mostly very localised, often small, and in KwaZulu-Natal, frequently limited to a few scattered, ... [details]

 Ecology

Known from a single collection made near Antsirabe in the Antananarivo province of Madagascar. The only known ... [details]

 Ecology

Rocky areas at high elevation (1,300–2,500 m). Flowering specimens were collected from December to April, ... [details]

 Ecology

The species is found infrequent on granitic formations in ericaceous shrubland, grassland and open forests at mid ... [details]

 Ecology

This taxon occurs in ericoid shrublands at (very) high elevations (1,400–2,800 m). Inflorescences are initiated ... [details]

 Ecology

This species is restricted to the herbaceous fringe of lower montane inselbergs. It has been observed from 500 to ... [details]

 Identification

1 Plants <30 cm tall with flowering culm scarcely exceeding the basal leaves; cauline leaves absent ..........2 1 Plants >30 cm tall with flowering culm exceeding the leaves; cauline leaves present ..........3 2 Spikelets ... [details]