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Cyperaceae taxon details

Carex divulsa Stokes

1676851  (urn:lsid:marinespecies.org:taxname:1676851)

accepted
Species
Carex canescens Huds. · unaccepted
Carex muricata Desf. · unaccepted
Carex nemorosa Lumn. · unaccepted
Carex persica Nelmes · unaccepted
Carex virens Lam. · unaccepted

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  1. Subspecies Carex divulsa subsp. leersiana Rauschert (uncertain > unassessed)
  2. Forma Carex divulsa f. angustifolia (Podp.) Soó accepted as Carex divulsa Stokes
  3. Forma Carex divulsa f. guestphalica (Rchb.) Kük. accepted as Carex divulsa Stokes
  4. Forma Carex divulsa f. misera (Kük. ex Vollm.) Kük. accepted as Carex divulsa Stokes
  5. Forma Carex divulsa f. polycarpa (Vollm.) Kük. accepted as Carex divulsa Stokes
  6. Subspecies Carex divulsa subsp. chabertii (F.W.Schultz) Asch. & Graebn. accepted as Carex leersii F.W.Schultz
  7. Subspecies Carex divulsa subsp. coriogyne (Nelmes) Ö.Nilsson accepted as Carex coriogyne Nelmes
  8. Subspecies Carex divulsa subsp. divulsa Stokes accepted as Carex divulsa Stokes
  9. Subspecies Carex divulsa subsp. leersii (Kneuck.) W.Koch accepted as Carex leersii F.W.Schultz
  10. Subspecies Carex divulsa subsp. orsiniana (Ten.) K.Richt. accepted as Carex muricata subsp. muricata L.
  11. Subspecies Carex divulsa subsp. virens (Lam.) Nyman accepted as Carex divulsa Stokes
  12. Variety Carex divulsa var. approximata Legrand accepted as Carex divulsa Stokes
  13. Variety Carex divulsa var. chabertii (F.W.Schultz) Nyman accepted as Carex leersii F.W.Schultz
  14. Variety Carex divulsa var. congesta Gren. accepted as Carex divulsa Stokes
  15. Variety Carex divulsa var. contigua (Hoppe) Nyman accepted as Carex spicata subsp. spicata
  16. Variety Carex divulsa var. guestphalica (Rchb.) F.W.Schultz ex Asch. & Graebn. accepted as Carex divulsa Stokes
  17. Variety Carex divulsa var. intermedia Lange accepted as Carex leersii F.W.Schultz
  18. Variety Carex divulsa var. javanica Nelmes accepted as Carex divulsa Stokes
  19. Variety Carex divulsa var. misera Kük. ex Vollm. accepted as Carex divulsa Stokes
  20. Variety Carex divulsa var. pairae (F.W.Schultz) Nyman accepted as Carex pairae F.W.Schultz
  21. Variety Carex divulsa var. polycarpa Vollm. accepted as Carex divulsa Stokes
  22. Variety Carex divulsa var. virens (Lam.) Steud. accepted as Carex divulsa Stokes
  23. Variety Carex divulsa var. virens (Lam.) Mérat accepted as Carex divulsa Stokes
  24. Variety Carex divulsa var. virens (Lam.) Steud. accepted as Carex divulsa Stokes
terrestrial
Withering, W.; Stokes, J. (1787). A botanical arrangement of British plants; including the uses of each species, in medicine, diet, rural economy and the arts, with an easy introduction to the study of botany ... /. , available online at https://www.biodiversitylibrary.org/page/56419277 [details] 
Holotype  K 000308960  
Holotype K 000308960 [details]
Description **Rizoma grueso, que forma céspedes hasta de 60(80) cm de diámetro. Tallos (15)25–60(90) cm, trígonos, con los...  
Description **Rizoma grueso, que forma céspedes hasta de 60(80) cm de diámetro. Tallos (15)25–60(90) cm, trígonos, con los ángulos agudos, escábridos en la parte superior. Hojas (1,5)2,5–3,5(4,3) mm de anchura, de menor longitud que los tallos, rara vez mayor, planas, algo ásperas en los bordes, no muy rígidas; lígula (0,3)1,5–2(3) mm, generalmente de longitud algo menor que la anchura, de ápice obtuso o redondeado; generalmente sin antelígula; vainas basales poco o muy fibrosas, de color pardo. Bráctea inferior de longitud mucho menor que la inflorescencia, setácea, no envainante. Espigas dispuestas en espiga simple de 2,3–17 cm, rara vez con alguna rama en la base. Glumas ovales, acuminadas a mucronadas, de color pardo oscuro, rara vez hialino-verdoso o pálido. Utrículos 3,2–5,5 × 1,5–2,2 mm, de contorno estrechamente oval y base ± atenuada, plano-convexos, con nerviación apenas perceptible o con un número variable de nervios bien marcados, verdosos, generalmente atenuados en un pico de 0,5–1,4 mm, escábrido-serrulado, bífido. Aquenios 1,9–2,5 × (1,2)1,7–2,3 mm, de contorno elíptico, oblongo u orbicular, biconvexos, rara vez plano-convexos, con la base del estilo persistente en forma de una corta columna. [details]

Description Caespitose 15-90 cm, forming small tufts, with short stolons. Stem sharply trigonous above, edges scabrous. Leaves soft,...  
Description Caespitose 15-90 cm, forming small tufts, with short stolons. Stem sharply trigonous above, edges scabrous. Leaves soft, from c. ½ to as long as stem; sheaths 15-40 mm, whitish to grey-brown or reddish, margin of scarious side concave or straight; ligule 0.1-0.2 mm, arch as high as wide or higher; blades 2-3.5 mm wide, flat, wrinkled along main veins, adaxial side smooth or +/- papillose, basal baxial side smooth , margins, apex and mid-vein scabrous. Inflorescence 30-70 mm, of 3-8(-15) androgynous spikes, overlapping or a few lowest distant, lowest branch sometimes elongate and with several spikes. Bracts sheathless, glume-like or at most as long as their spike, very narrow, green, scabrous. Spikes globular, 5-7 mm diam., green, later light brown; glumes 2.5-4.6 x 1.3-2.5 mm, ovate, obtuse to acute or frequently with mucro up to 0.5 mm, mid-vein white, surrounded by green area, bordered by two, white, conspicuous veins, margins scarious; utricles 3.3-5.4 x 1.5 mm, plano-convex, narrowly ovoid, narrowly winged. brownish, adaxial side without veins, glossy, baxial side without or with few veins, sometimes with short stipe 0.2-0.3 mm, beak green, winged, 1-2.3 mm, with two long, sharp teeth, ostiole white-margined. Nut 1.7-2.7 x 1.4-1.9 mm, bi-convex or planoconvex, widely ovoid to ellipsoid, glossy, finely verrucose. [details]
Cyperaceae Working Group. (2025). [see How to cite]. Global Cyperaceae Database. Carex divulsa Stokes. Accessed at: https://cyperaceae.org/aphia.php?p=taxdetails&id=1676851 on 2026-05-12
Date
action
by
2023-09-06 07:28:28Z
created
2024-12-10 11:47:54Z
unchecked
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2025-03-11 11:19:03Z
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Nomenclature

original description Withering, W.; Stokes, J. (1787). A botanical arrangement of British plants; including the uses of each species, in medicine, diet, rural economy and the arts, with an easy introduction to the study of botany ... /. , available online at https://www.biodiversitylibrary.org/page/56419277 [details] 

Taxonomy

taxonomy source Jermy, A.C.; Simpson, D.A.; Foley, M.J.Y.; Porter, M.S. (2007). Sedges of the British Isles. 554 pp. Botanical Society of Britain & Ireland, United Kingdom. [details] Available for editors  PDF available [request]

taxonomy source Jiménez-Mejías, P.; Martín-Bravo, S.; Márquez-Corro, J. I.; Donadío, S.; Roalson, E. H.; Naczi, R. F. C. (2021). A synopsis of the androgynous species of <i>Carex</i> subgenus <i>Vignea</i> (Cyperaceae) in South America. <em>Botanical Journal of the Linnean Society.</em> 196(2): 188-220., available online at https://doi.org/10.1093/botlinnean/boaa100 [details] OpenAccess publication

taxonomy source Luceño, M., Castroviejo, S., Jiménez-Mejías, P. (2008). Cyperaceae. In: Castroviejo, Luceño, M., Galán, A., Jiménez-Mejías, P., Cabezas, F., Medina, L. (eds). Flora iberica. Plantas vasculares de la Península Ibérica e Islas Baleares. Vol. XVIII. Madrid: CSIC., available online at https://bibdigital.rjb.csic.es/records/item/12958-flora-iberica-vol-18-cyperaceae-pontederiaceae?offset=, http://www.floraiberica.es/PHP/familias_lista_.php?familia=Cyperaceae [details] 

Other

subsequent type designation Molina, A.; Acedo, C.; Llamas, F. (2008). Taxonomy and new taxa of the Carex divulsa aggregate in Eurasia (section Phaestoglochin, Cyperaceae). <em>Botanical Journal of the Linnean Society.</em> 156(3): 385-409., available online at https://doi.org/10.1111/j.1095-8339.2007.00760.x
page(s): 395 [details] Available for editors  PDF available [request]

biology source Jiménez-Mejías, P.; Fabbroni, M.; Donadío, S.; Rodríguez-Palacios, G. E.; Hilpold, A.; Martín-Bravo, S.; Waterway, M. J.; Roalson, E. H. (2016). Taxonomic and distribution notes on Carex (Cyperaceae) from the Neotropics. Boletin de la Sociedad Argentina de Botanica, 51(4): 727-739., available online at https://doi.org/10.31055/1851.2372.v51.n4.16400 [details] Available for editors  PDF available [request]

 
 Present  Inaccurate  Introduced: alien  Containing type locality 
   

Holotype K 000308960 [details]
From editor or global species database
Description **Rizoma grueso, que forma céspedes hasta de 60(80) cm de diámetro. Tallos (15)25–60(90) cm, trígonos, con los ángulos agudos, escábridos en la parte superior. Hojas (1,5)2,5–3,5(4,3) mm de anchura, de menor longitud que los tallos, rara vez mayor, planas, algo ásperas en los bordes, no muy rígidas; lígula (0,3)1,5–2(3) mm, generalmente de longitud algo menor que la anchura, de ápice obtuso o redondeado; generalmente sin antelígula; vainas basales poco o muy fibrosas, de color pardo. Bráctea inferior de longitud mucho menor que la inflorescencia, setácea, no envainante. Espigas dispuestas en espiga simple de 2,3–17 cm, rara vez con alguna rama en la base. Glumas ovales, acuminadas a mucronadas, de color pardo oscuro, rara vez hialino-verdoso o pálido. Utrículos 3,2–5,5 × 1,5–2,2 mm, de contorno estrechamente oval y base ± atenuada, plano-convexos, con nerviación apenas perceptible o con un número variable de nervios bien marcados, verdosos, generalmente atenuados en un pico de 0,5–1,4 mm, escábrido-serrulado, bífido. Aquenios 1,9–2,5 × (1,2)1,7–2,3 mm, de contorno elíptico, oblongo u orbicular, biconvexos, rara vez plano-convexos, con la base del estilo persistente en forma de una corta columna. [details]

Description Caespitose 15-90 cm, forming small tufts, with short stolons. Stem sharply trigonous above, edges scabrous. Leaves soft, from c. ½ to as long as stem; sheaths 15-40 mm, whitish to grey-brown or reddish, margin of scarious side concave or straight; ligule 0.1-0.2 mm, arch as high as wide or higher; blades 2-3.5 mm wide, flat, wrinkled along main veins, adaxial side smooth or +/- papillose, basal baxial side smooth , margins, apex and mid-vein scabrous. Inflorescence 30-70 mm, of 3-8(-15) androgynous spikes, overlapping or a few lowest distant, lowest branch sometimes elongate and with several spikes. Bracts sheathless, glume-like or at most as long as their spike, very narrow, green, scabrous. Spikes globular, 5-7 mm diam., green, later light brown; glumes 2.5-4.6 x 1.3-2.5 mm, ovate, obtuse to acute or frequently with mucro up to 0.5 mm, mid-vein white, surrounded by green area, bordered by two, white, conspicuous veins, margins scarious; utricles 3.3-5.4 x 1.5 mm, plano-convex, narrowly ovoid, narrowly winged. brownish, adaxial side without veins, glossy, baxial side without or with few veins, sometimes with short stipe 0.2-0.3 mm, beak green, winged, 1-2.3 mm, with two long, sharp teeth, ostiole white-margined. Nut 1.7-2.7 x 1.4-1.9 mm, bi-convex or planoconvex, widely ovoid to ellipsoid, glossy, finely verrucose. [details]

Translation Rhizome thick, forming tussocks up to 60(80) cm in diameter. Stems (15)25–60(90) cm, trigonous, with acute angles, scabrid in the upper part. Leaves (1.5)2.5–3.5(4.3) mm wide, shorter than the stems, rarely longer, flat, somewhat rough on the margins, not very rigid; ligule (0.3)1.5–2(3) mm, generally slightly shorter than wide, with an obtuse or rounded apex; anteligule generally absent; basal sheaths little to strongly fibrous, brown. Lowest bract much shorter than the inflorescence, setaceous, not sheathing. Spikes arranged in a simple spike 2.3–17 cm long, rarely with a branch at the base. Glumes ovate, acuminate to mucronate, dark brown, rarely hyaline-green or pale. Utricles 3.2–5.5 × 1.5–2.2 mm, narrowly ovate in outline, base ± attenuate, plano-convex, with venation scarcely visible or with a variable number of well-marked nerves, greenish, generally attenuate into a beak 0.5–1.4 mm long, scabrid-serrulate, bifid. Achenes 1.9–2.5 × (1.2)1.7–2.3 mm, elliptic, oblong or orbicular in outline, biconvex, rarely plano-convex, with the persistent style base forming a short column. [details]
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