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Cyperaceae taxon details

Carex atrata L.

To Royal Botanic Gardens, Kew (Carex atrata K000998994 type 1)
To Royal Botanic Gardens, Kew (Carex atrata K000998994 type 1)

1677257  (urn:lsid:marinespecies.org:taxname:1677257)

  1. Biota
  2. Plantae (Kingdom)
  3. Viridiplantae (Subkingdom)
  4. Streptophyta (Infrakingdom)
  5. Tracheophyta (Phylum (Division))
  6. Spermatophytina (Subphylum (Subdivision))
  7. Magnoliopsida (Class)
  8. Lilianae (Superorder)
  9. Poales (Order)
  10. Cyperaceae (Family)
  11. Cariceae (Tribe)
  12. Carex (Genus)
  13. Carex subg. Carex (Subgenus)
  14. Carex sect. Racemosae (Section)
  15. Carex atrata (Species)
accepted
Species

Ordering

  • Alphabetically
  • By status

Children Display

  1. Subspecies Carex atrata subsp. atrata
  2. Subspecies Carex atrata subsp. longistolonifera (Kük.) S.Yun Liang
  3. Subspecies Carex atrata subsp. pullata (Boott) Kük.
  4. Subspecies Carex atrata subsp. aterrima (Hoppe) Čelak (uncertain > unassessed)
  5. Subspecies Carex atrata subsp. aterrima (Hoppe) S.Yun Liang (uncertain > unassessed)
  6. Variety Carex atrata var. nigra Boott (uncertain > unassessed)
  7. Forma Carex atrata f. aterrima (Hoppe) Neuman accepted as Carex aterrima Hoppe
  8. Forma Carex atrata f. caeca (L.H.Bailey) Kük. accepted as Carex heteroneura var. heteroneura
  9. Forma Carex atrata f. castanea (Nyman) K.Richt. accepted as Carex atrata subsp. atrata
  10. Forma Carex atrata f. decolorans Neuman accepted as Carex atrata subsp. atrata
  11. Forma Carex atrata f. gelida Schur accepted as Carex atrata subsp. atrata
  12. Forma Carex atrata f. lenis (L.H.Bailey) Kük. accepted as Carex heteroneura var. heteroneura
  13. Forma Carex atrata f. parviflora (Host) Neuman accepted as Carex parviflora Host
  14. Forma Carex atrata f. wolfii (Kneuck.) Kük. accepted as Carex aterrima subsp. aterrima
  15. Subforma Carex atrata subforma aristata Podp. accepted as Carex atrata subsp. atrata
  16. Subspecies Carex atrata subsp. apodostachya (Ohwi) T.Koyama accepted as Carex atrata subsp. atrata
  17. Subspecies Carex atrata subsp. aterrima (Hoppe) Hartm. accepted as Carex aterrima Hoppe
  18. Subspecies Carex atrata subsp. atratiformis Kük. accepted as Carex atratiformis Britton
  19. Subspecies Carex atrata subsp. atrosquama (Mack.) Hultén accepted as Carex atrosquama Mack.
  20. Subspecies Carex atrata subsp. bicolor Čelak. accepted as Carex atrata subsp. atrata
  21. Subspecies Carex atrata subsp. castanea (Nyman) K.Richt. accepted as Carex atrata subsp. atrata
  22. Subspecies Carex atrata subsp. caucasica (Steven) Kük. accepted as Carex caucasica Steven
  23. Subspecies Carex atrata subsp. magellanica (Lam.) Wahlenb. accepted as Carex magellanica Lam.
  24. Subspecies Carex atrata subsp. nigra (Gaudin) Hartm. accepted as Carex parviflora Host
  25. Subspecies Carex atrata subsp. perfusca (V.I.Krecz.) T.Koyama accepted as Carex aterrima subsp. aterrima
  26. Variety Carex atrata var. aterrima (Hoppe) Winckler accepted as Carex aterrima Hoppe
  27. Variety Carex atrata var. atrata accepted as Carex atrata L.
  28. Variety Carex atrata var. atrosquama (Mack.) Cronquist accepted as Carex atrosquama Mack.
  29. Variety Carex atrata var. bicolor Čelak. accepted as Carex atrata subsp. atrata
  30. Variety Carex atrata var. brunnescens Andersson accepted as Carex atrata subsp. atrata
  31. Variety Carex atrata var. castanea Nyman accepted as Carex atrata subsp. atrata
  32. Variety Carex atrata var. caucasica (Steven) Boott accepted as Carex caucasica Steven
  33. Variety Carex atrata var. chalciolepis (Holm) Kük. accepted as Carex chalciolepis Holm
  34. Variety Carex atrata var. conglomerata Neilr. accepted as Carex parviflora Host
  35. Variety Carex atrata var. discolor L.H.Bailey accepted as Carex bella L.H.Bailey
  36. Variety Carex atrata var. dubia Gaudin accepted as Carex atrata subsp. atrata
  37. Variety Carex atrata var. erecta W.Boott accepted as Carex heteroneura var. heteroneura
  38. Variety Carex atrata var. glacialis Boott accepted as Carex atrata subsp. pullata (Boott) Kük.
  39. Variety Carex atrata var. heterostachya Almq. accepted as Carex atrata subsp. atrata
  40. Variety Carex atrata var. japonoalpina T.Koyama accepted as Carex atrata subsp. atrata
  41. Variety Carex atrata var. larimerana (Kelso) Kelso accepted as Carex chalciolepis Holm
  42. Variety Carex atrata var. laxa Neilr. accepted as Carex atrata subsp. atrata
  43. Variety Carex atrata var. longistolonifera Kük. accepted as Carex atrata subsp. longistolonifera (Kük.) S.Yun Liang
  44. Variety Carex atrata var. magellanica (Lam.) Wahlenb. accepted as Carex magellanica Lam.
  45. Variety Carex atrata var. nigra Olney accepted as Carex nova L.H.Bailey
  46. Variety Carex atrata var. nigra Gaudin accepted as Carex parviflora Host
  47. Variety Carex atrata var. nigra (All.) W.Boott accepted as Carex helleri Mack.
  48. Variety Carex atrata var. ovata Boott accepted as Carex atrata subsp. atrata
  49. Variety Carex atrata var. parviflora (Host) Beck accepted as Carex parviflora Host
  50. Variety Carex atrata var. pseudocilicica Kük. accepted as Carex atrata subsp. atrata
  51. Variety Carex atrata var. pullata Boott accepted as Carex atrata subsp. pullata (Boott) Kük.
  52. Variety Carex atrata var. rectiuscula Hartm. accepted as Carex atrata subsp. atrata
  53. Variety Carex atrata var. sinensis Kük. accepted as Carex atrata subsp. pullata (Boott) Kük.
  54. Variety Carex atrata var. spadicea Beurl. accepted as Carex atrata subsp. atrata
  55. Variety Carex atrata var. spiculosior Norman accepted as Carex atrata subsp. atrata
  56. Variety Carex atrata var. subgracilenta Kük. accepted as Carex atrata subsp. pullata (Boott) Kük.
  57. Variety Carex atrata var. typica Beck accepted as Carex atrata L.
  58. Variety Carex atrata var. varia Gaudin accepted as Carex atrata L.
terrestrial
Linnaeus, C. (1753). Species plantarum, exhibentes plantas rite cognitas ad genera relatas cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas. Vol 2. Salvius, Stockholm., available online at https://www.biodiversitylibrary.org/page/358580
page(s): 257 [details] 
Lectotype  LINN HL1100-52, geounit Sweden  
Lectotype LINN HL1100-52, geounit Sweden [details]
Description Rhizomes short; shoots tufted; roots pale brown; scales brownish or reddish-purple, persistent. Stems (25–)30–55 cm,...  
Description Rhizomes short; shoots tufted; roots pale brown; scales brownish or reddish-purple, persistent. Stems (25–)30–55 cm, thick, rigid, trigonous, solid. Leaves 15–30 cm × 2–6 mm, ± flat or slightly keeled, abruptly tapered to a rough apex, mid green to deep blue-green; hypostomous; sheaths dark brown, often wine-red, persistent, inner face hyaline, persistent; apex straight; ligule 2–3 mm, obtuse to acute. Inflorescence 1/8–1/6 length of stem; bracts leaf-like, lower exceeding inflorescence, lowest not or scarcely sheathing. Spikes 3–5, ovoid, 8–20 mm, clustered in an inclined head, becoming ± nodding in fruit, terminal spike female at top, male at base, lower female only; lower peduncles rough, as long as spike. Male glumes 4.5–6 mm, ovate-elliptic, red-black with pale midrib; apex acute or rounded. Female glumes 3.5–4.5 mm, ovate-elliptic, purple or reddish-black with pale midrib; apex ± acute. Utricles 3–4 mm, obovoid-ellipsoid, compressed at edges, green; beak 0.3–0.5 mm, slightly notched; stigmas 3; nut obovoid, trigonous. [details]

Description Plants loosely cespitose. Culms (10–)30–40(–60) cm, smooth. Leaves 3–5 mm wide. Inflorescences: proximal bracts...  
Description Plants loosely cespitose. Culms (10–)30–40(–60) cm, smooth. Leaves 3–5 mm wide. Inflorescences: proximal bracts shorter than or exceeding inflorescences; spikes overlapping, erect or proximal spikes sometimes spreading, distinct, contiguous, short-pendunculate, short-oblong or elongate, 10–25 × 6–10 mm; lateral 2–4(–6) spikes pistillate, pedunculate, of similar length; terminal spike gynecandrous. Pistillate scales dark brown or black to margins, lanceolate, longer than and as broad as perigynia, midvein same color as body, inconspicuous or lighter in color and conspicuous, acute. Perigynia ascending, golden or dark brown, veinless, ovate, 3–4 × 1.5–1.75 mm, apex abruptly beaked, distally papillose; beak 04–0.5 mm, bidentate, smooth. Achenes filling proximal 1/2 or less of perigynia. 2n = 54. [details]

Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 976 1753 to Sp. Pl. : 976 (1753), information provided by...  
Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 976 1753 to Sp. Pl. : 976 (1753), information provided by Alan E. on email Jun. 07 2021 More details could be found in The Plant L [details]
Cyperaceae Working Group. (2025). [see How to cite]. Global Cyperaceae Database. Carex atrata L.. Accessed at: https://cyperaceae.org/aphia.php?p=taxdetails&id=1677257 on 2026-05-31
Date
action
by
2023-09-06 07:28:28Z
created
Jiménez-Mejías, Pedro
2024-12-10 11:47:54Z
unchecked
db_admin
2025-03-11 11:19:03Z
changed
Márquez-Corro, José Ignacio

Creative Commons License The webpage text is licensed under a Creative Commons Attribution 4.0 License


Nomenclature

original description Linnaeus, C. (1753). Species plantarum, exhibentes plantas rite cognitas ad genera relatas cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas. Vol 2. Salvius, Stockholm., available online at https://www.biodiversitylibrary.org/page/358580
page(s): 257 [details] 

basis of record Plants of the World Online (POWO). , available online at https://powo.science.kew.org/ [details] 

Taxonomy

taxonomy source Jermy, A.C.; Simpson, D.A.; Foley, M.J.Y.; Porter, M.S. (2007). Sedges of the British Isles. 554 pp. Botanical Society of Britain & Ireland, United Kingdom. [details] Available for editors  PDF available [request]

Other

additional source Li, B. Z.; Zhang, S. R. (2024). An updated species checklist and taxonomic synopsis of Cyperaceae in China. <em>Biodiversity Science.</em> 32(7): 24106., available online at https://doi.org/10.17520/biods.2024106 [details] Available for editors  PDF available [request]

 
 Present  Inaccurate  Introduced: alien  Containing type locality 
   

Lectotype LINN HL1100-52, geounit Sweden [details]
Syntype K 000998994, geounit India [details]
From editor or global species database
Additional information 2n=56 [details]

Additional information 2n=52, 54 [details]

Description Rhizomes short; shoots tufted; roots pale brown; scales brownish or reddish-purple, persistent. Stems (25–)30–55 cm, thick, rigid, trigonous, solid. Leaves 15–30 cm × 2–6 mm, ± flat or slightly keeled, abruptly tapered to a rough apex, mid green to deep blue-green; hypostomous; sheaths dark brown, often wine-red, persistent, inner face hyaline, persistent; apex straight; ligule 2–3 mm, obtuse to acute. Inflorescence 1/8–1/6 length of stem; bracts leaf-like, lower exceeding inflorescence, lowest not or scarcely sheathing. Spikes 3–5, ovoid, 8–20 mm, clustered in an inclined head, becoming ± nodding in fruit, terminal spike female at top, male at base, lower female only; lower peduncles rough, as long as spike. Male glumes 4.5–6 mm, ovate-elliptic, red-black with pale midrib; apex acute or rounded. Female glumes 3.5–4.5 mm, ovate-elliptic, purple or reddish-black with pale midrib; apex ± acute. Utricles 3–4 mm, obovoid-ellipsoid, compressed at edges, green; beak 0.3–0.5 mm, slightly notched; stigmas 3; nut obovoid, trigonous. [details]

Description Plants loosely cespitose. Culms (10–)30–40(–60) cm, smooth. Leaves 3–5 mm wide. Inflorescences: proximal bracts shorter than or exceeding inflorescences; spikes overlapping, erect or proximal spikes sometimes spreading, distinct, contiguous, short-pendunculate, short-oblong or elongate, 10–25 × 6–10 mm; lateral 2–4(–6) spikes pistillate, pedunculate, of similar length; terminal spike gynecandrous. Pistillate scales dark brown or black to margins, lanceolate, longer than and as broad as perigynia, midvein same color as body, inconspicuous or lighter in color and conspicuous, acute. Perigynia ascending, golden or dark brown, veinless, ovate, 3–4 × 1.5–1.75 mm, apex abruptly beaked, distally papillose; beak 04–0.5 mm, bidentate, smooth. Achenes filling proximal 1/2 or less of perigynia. 2n = 54. [details]

Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 976 1753 to Sp. Pl. : 976 (1753), information provided by Alan E. on email Jun. 07 2021 More details could be found in The Plant L [details]
    Definitions

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LanguageName 
Japanese クロボスゲ（Kurobo-suge）  [details]
To Biodiversity Heritage Library (1 publication) (from synonym Loxanisa atrata (L.) Raf.)
To Biodiversity Heritage Library (1 publication) (from synonym Carex atrata var. typica Beck)
To Biodiversity Heritage Library (4 publications) (from synonym Trasus atratus (L.) Gray)
To Biodiversity Heritage Library (729 publications) 
To European Nucleotide Archive, ENA (Carex atrata) 
To International Plant Names Index (IPNI) 
To International Plant Names Index (IPNI) (from synonym Loxanisa atrata (L.) Raf.)
To International Plant Names Index (IPNI) (from synonym Trasus atratus (L.) Gray)
To International Plant Names Index (IPNI) (from synonym Carex atrata var. varia Gaudin)
To International Plant Names Index (IPNI) (from synonym Carex atrata var. typica Beck)
To Plants of the World Online (from synonym Carex atrata var. varia Gaudin)
To Plants of the World Online (from synonym Carex atrata var. typica Beck)
To Plants of the World Online 
To Plants of the World Online (from synonym Loxanisa atrata (L.) Raf.)
To Plants of the World Online (from synonym Trasus atratus (L.) Gray)
To Royal Botanic Gardens, Kew (Carex atrata K000998994 type 1) 
To The Plant List (from synonym Carex atrata var. atrata)
From editor or global species database
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