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Cyperaceae taxon details

Carex vesicaria L.

1677354  (urn:lsid:marinespecies.org:taxname:1677354)

accepted
Species
Carex furcata Lapeyr. · unaccepted
Carex monile Tuck. · unaccepted
Carex pendula Moench · unaccepted
Carex raeana Boott · unaccepted
Carex suilla J.Fellm. · unaccepted
Carex vaseyi Dewey · unaccepted
Carex vesiciformis Peterm. · unaccepted > homotypic synonym

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  1. Variety Carex vesicaria var. physocarpa Kük. (uncertain > unassessed)
  2. Variety Carex vesicaria var. saxatilis (L.) H.Lindb. (uncertain > unassessed)
  3. Forma Carex vesicaria f. alpigena (Fr.) Neuman accepted as Carex vesicaria L.
  4. Forma Carex vesicaria f. aurea Lange accepted as Carex vesicaria L.
  5. Forma Carex vesicaria f. brachystachys Lindeb. accepted as Carex vesicaria L.
  6. Forma Carex vesicaria f. dichroa (Andersson) Kük. accepted as Carex vesicaria L.
  7. Forma Carex vesicaria f. dichroa (Andersson) Neuman accepted as Carex vesicaria L.
  8. Forma Carex vesicaria f. elatior Andersson accepted as Carex vesicaria L.
  9. Forma Carex vesicaria f. fluitans Raymond accepted as Carex vesicaria L.
  10. Forma Carex vesicaria f. globosa (Olney) Kük. accepted as Carex vesicaria L.
  11. Forma Carex vesicaria f. gracillima Kük. accepted as Carex vesicaria L.
  12. Forma Carex vesicaria f. hexasticha Hoschedé accepted as Carex vesicaria L.
  13. Forma Carex vesicaria f. laxa (Trautv.) Kük. accepted as Carex saxatilis L.
  14. Forma Carex vesicaria f. monstrosa (L.H.Bailey) Kük. accepted as Carex vesicaria L.
  15. Forma Carex vesicaria f. pendulina Blytt accepted as Carex vesicaria L.
  16. Forma Carex vesicaria f. polystachya (Blytt) Kük. accepted as Carex vesicaria L.
  17. Forma Carex vesicaria f. pungens (L.H.Bailey) Kük. accepted as Carex vesicaria L.
  18. Forma Carex vesicaria f. tenuistachya (Kük.) T.Koyama accepted as Carex vesicata Meinsh.
  19. Forma Carex vesicaria f. umbrosa Kük. accepted as Carex vesicaria L.
  20. Subspecies Carex vesicaria subsp. alpigena (Fr.) Printz accepted as Carex vesicaria L.
  21. Subspecies Carex vesicaria subsp. dichroa Freyn accepted as Carex pamirensis subsp. dichroa Malyschev
  22. Subspecies Carex vesicaria subsp. flavescens Ehrh. accepted as Carex vesicaria L.
  23. Subspecies Carex vesicaria subsp. friesii (Blytt) Nyman accepted as Carex friesii Blytt represented as Carex ×friesii Blytt
  24. Subspecies Carex vesicaria subsp. lacustris Th.Fr. accepted as Carex utriculata Boott
  25. Subspecies Carex vesicaria subsp. plumbea Wahlenb. accepted as Carex riparia Curtis
  26. Subspecies Carex vesicaria subsp. pulla (Gooden.) Andersson accepted as Carex saxatilis L.
  27. Subspecies Carex vesicaria subsp. saxatilis (L.) Kük. accepted as Carex saxatilis L.
  28. Subspecies Carex vesicaria subsp. saxatilis (L.) C.Hartm. accepted as Carex saxatilis L.
  29. Subspecies Carex vesicaria subsp. subfusca Ehrh. accepted as Carex vesicaria L.
  30. Subspecies Carex vesicaria subsp. sylvatica Ehrh. accepted as Carex sylvatica subsp. sylvatica
  31. Subspecies Carex vesicaria subsp. vesicata (Meinsh.) T.V.Egorova accepted as Carex vesicata Meinsh.
  32. Subspecies Carex vesicaria subsp. virens F.Nyl. accepted as Carex vesicaria L.
  33. Variety Carex vesicaria var. alpigena Fr. accepted as Carex vesicaria L.
  34. Variety Carex vesicaria var. alpigena Kük. accepted as Carex pulla subsp. dichroa Freyn accepted as Carex pamirensis subsp. dichroa Malyschev
  35. Variety Carex vesicaria var. brachystachys Blytt accepted as Carex vesicaria L.
  36. Variety Carex vesicaria var. calcifoenum Laest. accepted as Carex vesicaria L.
  37. Variety Carex vesicaria var. colorata (L.H.Bailey) Kük. accepted as Carex vesicaria L.
  38. Variety Carex vesicaria var. compacta Dewey accepted as Carex membranacea Hook.
  39. Variety Carex vesicaria var. compacta (R.Br.) Kük. accepted as Carex membranacea Hook.
  40. Variety Carex vesicaria var. dichroa Andersson accepted as Carex vesicaria L.
  41. Variety Carex vesicaria var. distenta Fr. accepted as Carex saxatilis L.
  42. Variety Carex vesicaria var. fatua Peterm. accepted as Carex vesicaria L.
  43. Variety Carex vesicaria var. friesii (Blytt) Kük. accepted as Carex friesii Blytt represented as Carex ×friesii Blytt
  44. Variety Carex vesicaria var. globosa Olney accepted as Carex vesicaria L.
  45. Variety Carex vesicaria var. grahamii (Boott) Kük. accepted as Carex grahamii Boott represented as Carex ×grahamii Boott
  46. Variety Carex vesicaria var. inalpina Laest. accepted as Carex rostrata var. rostrata
  47. Variety Carex vesicaria var. involuta Bab. accepted as Carex involuta (Bab.) Syme represented as Carex ×involuta (Bab.) Syme
  48. Variety Carex vesicaria var. jejuna Fernald accepted as Carex saxatilis L.
  49. Variety Carex vesicaria var. lanceolata Olney accepted as Carex vesicaria L.
  50. Variety Carex vesicaria var. latifolia Blytt accepted as Carex vesicaria L.
  51. Variety Carex vesicaria var. laurentiana Fernald accepted as Carex saxatilis L.
  52. Variety Carex vesicaria var. major Boott accepted as Carex exsiccata L.H.Bailey
  53. Variety Carex vesicaria var. monile (Tuck.) Boeckeler accepted as Carex vesicaria L.
  54. Variety Carex vesicaria var. monile (Tuck. ex Boott) Fernald accepted as Carex vesicaria L.
  55. Variety Carex vesicaria var. monstrosa (L.H.Bailey) Farw. accepted as Carex vesicaria L.
  56. Variety Carex vesicaria var. obtusisquamis L.H.Bailey accepted as Carex vesicaria L.
  57. Variety Carex vesicaria var. pacifica (L.H.Bailey) Kük. accepted as Carex vesicaria L.
  58. Variety Carex vesicaria var. pamirica O.Fedtsch. accepted as Carex pamirensis C.B.Clarke
  59. Variety Carex vesicaria var. physocarpa (C.Presl) Kük. accepted as Carex saxatilis L.
  60. Variety Carex vesicaria var. plumbea Wahlenb. accepted as Carex riparia Curtis
  61. Variety Carex vesicaria var. raeana (Boott) Fernald accepted as Carex vesicaria L.
  62. Variety Carex vesicaria var. reflexa Meinsh. accepted as Carex mollissima Christ ex Scheutz
  63. Variety Carex vesicaria var. robusta Sond. accepted as Carex vesicaria L.
  64. Variety Carex vesicaria var. rostratiformis Waisb. accepted as Carex involuta (Bab.) Syme represented as Carex ×involuta (Bab.) Syme
  65. Variety Carex vesicaria var. subalpina Laest. accepted as Carex vesicaria L.
  66. Variety Carex vesicaria var. tenuistachya Kük. accepted as Carex vesicata Meinsh.
  67. Variety Carex vesicaria var. utriculata (Boott) Dewey accepted as Carex utriculata Boott
  68. Variety Carex vesicaria var. vesicaria accepted as Carex vesicaria L.
  69. Variety Carex vesicaria var. virens (F.Nyl.) Nyl. & Saelán accepted as Carex vesicaria L.
marine, brackish, fresh, terrestrial
recent only
Linnaeus, C. (1753). Species plantarum, exhibentes plantas rite cognitas ad genera relatas cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas. Vol 2. Salvius, Stockholm., available online at https://www.biodiversitylibrary.org/page/358580
page(s): 979 [details] 
Description Rhizomes shortly creeping; shoots slender, markedly trigonous, 2–3 in each tuft; roots thick, pale yellow-brown; scales...  
Description Rhizomes shortly creeping; shoots slender, markedly trigonous, 2–3 in each tuft; roots thick, pale yellow-brown; scales brown- or purple-red, usually persistent. Stems 30–120 cm, trigonous, but with a sharp, low ridge on the angles rough above, smooth below. Leaves up to 150 cm × 4–8 mm, serrulate at least towards tip, thin, rigid, plicate, gradually tapered to a fine point, mid green or yellow-green, quickly decaying; hypostomous; sheaths becoming purple-red, persistent, inner face hyaline, fibrillose on splitting, apex straight or concave; ligule 6–12 mm, acute. Inflorescence 1/4–1/3 length of stem; bracts leaf-like, lower exceeding inflorescence. Male spikes 2–4, 10–40 mm, the lower often with setaceous bracts; male glumes 4–6 mm, elliptic or oblanceolate, purplish-brown with green or paler midrib and hyaline margins; apex ± acute. Female spikes 2–3, ± contiguous but distant from the male spikes, 2–4(–7) cm, oblong-cylindric, erect, the lowest often pendent; female glumes 4–6 mm, narrowly lanceolate, purplish-brown with paler or green midrib; apex acute or acuminate, hyaline. Utricles 6–8 mm, ovoid-ellipsoid, inflated, ribbed, ± shiny, olive green to yellow-green, ascending; beak c. 2 mm, smooth, broadly notched; stigmas 3; nut obovoid, trigonous. [details]

Description Laxamente cespitosa, con rizoma robusto, de entrenudos algo largos. Tallos fértiles (20)35–100(110) cm, muy ásperos...  
Description Laxamente cespitosa, con rizoma robusto, de entrenudos algo largos. Tallos fértiles (20)35–100(110) cm, muy ásperos hacia el ápice, excepcionalmente lisos o casi cuadrangulares. Hojas (3)4–6(7) mm de anchura, de mayor longitud que los tallos, planas, muy ásperas en los bordes y nervio medio por el envés, ± rígidas; lígula (2)4–8 mm, de ápice obtuso; sin antelígula; vainas basales con limbo desarrollado, solo las más inferiores escuamiformes, de color pardo, raramente manchadas de color púrpura, enteras. Bráctea inferior folíacea, de mayor longitud que la inflorescencia, con frecuencia envainante. Espigas masculinas (1)2–3, de (8)20–40(55) mm, a veces andróginas, de lineares a estrechamente fusiformes; espigas femeninas 2–3, de (10)20–40(50) mm, generalmente cilíndricas, densifloras, la superior frecuentemente andrógina, distantes o las superiores aproximadas, pedunculadas, la inferior a veces colgante, el resto erectas. Glumas masculinas lineares a lanceoladas, agudas o subagudas, pardas, frecuentemente con margen escarioso en la zona apical; glumas femeninas lanceoladas a oblongas, obtusas, pardas o, excepcionalmente, negras, de longitud mucho mayor que los utrículos, de color pardo-rojizo, las de la zona media casi siempre con margen escarioso. Utrículos (3,9)4–6(8) × 1,2–3,2 mm, suberectos, ovoides o elipsoidales, inflados, pardo-amarillentos, brillantes, con los nervios prominentes, gradualmente atenuados en un pico de (1)1,1–2,2 mm, profundamente bífido, con el sinus ± agudo, liso. Aquenios 1,9–2,1 × 1–1,5 mm, de contorno oblongo o elíptico, trígonos, de color pardo. 2n = 74*, 82, 86*. [details]

Description Perennials. rhizome stoloniferous, stolons branched. Stems 30–100 cm tall, stout, triquetrous, scabrous on upper part....  
Description Perennials. rhizome stoloniferous, stolons branched. Stems 30–100 cm tall, stout, triquetrous, scabrous on upper part. Leaves shorter than stem, 3–8 mm wide, pale green, scabrous. Basal sheaths reddish brown, ladder-like fibrillose. Involucral bracts leaflike, lowermost shortly sheathed, upper not sheathed. Inflorescences spikes 4–6; terminal 2 or 3 spikes male, approximate, linear cylindric, 2–3.5 cm long, subsessile; remaining spikes female, oblong or oblong-cylindric, 3–7 cm long, densely many flowered, shortly pedunculate. Staminate scales brown, apex obtuse to acute. Pistillate scales pale brown, midrib green, apex acute or acuminate. Perigynia yellow-green, obliquely patent, longer than pistillate scales, ovoid, inflated trigonous, 6–8 mm long, several veined, glabrous, long beaked, mouth bidentate. Achenes obovoid, trigonous, 1.7–2 mm long. Pistil style slender, flexuous; stigmas 3. [details]

Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 979 1753 to Sp. Pl. : 979 (1753), information provided by...  
Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 979 1753 to Sp. Pl. : 979 (1753), information provided by Alan E. on email Jun. 07 2021 More details could be found in The Plant L [details]
Cyperaceae Working Group. (2025). [see How to cite]. Global Cyperaceae Database. Carex vesicaria L.. Accessed at: https://cyperaceae.org/aphia.php?p=taxdetails&id=1677354 on 2026-05-28
Date
action
by
2023-09-06 07:28:28Z
created
2024-12-10 11:47:54Z
unchecked
db_admin
2025-03-11 11:19:03Z
changed
2025-04-02 20:29:58Z
changed

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Nomenclature

original description Linnaeus, C. (1753). Species plantarum, exhibentes plantas rite cognitas ad genera relatas cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas. Vol 2. Salvius, Stockholm., available online at https://www.biodiversitylibrary.org/page/358580
page(s): 979 [details] 

original description (of Carex vesiciformis Peterm.) Petermann, W. L. (1844). Beiträge zur deutschen Flora. <em>Flora.</em> 27: 225-344., available online at https://www.biodiversitylibrary.org/page/39342
page(s): 339 [details] 

basis of record Plants of the World Online (POWO). , available online at https://powo.science.kew.org/ [details] 

Taxonomy

taxonomy source Jermy, A.C.; Simpson, D.A.; Foley, M.J.Y.; Porter, M.S. (2007). Sedges of the British Isles. 554 pp. Botanical Society of Britain & Ireland, United Kingdom. [details] Available for editors  PDF available [request]

taxonomy source Luceño, M., Castroviejo, S., Jiménez-Mejías, P. (2008). Cyperaceae. In: Castroviejo, Luceño, M., Galán, A., Jiménez-Mejías, P., Cabezas, F., Medina, L. (eds). Flora iberica. Plantas vasculares de la Península Ibérica e Islas Baleares. Vol. XVIII. Madrid: CSIC., available online at https://bibdigital.rjb.csic.es/records/item/12958-flora-iberica-vol-18-cyperaceae-pontederiaceae?offset=, http://www.floraiberica.es/PHP/familias_lista_.php?familia=Cyperaceae [details] 

Other

additional source Li, B. Z.; Zhang, S. R. (2024). An updated species checklist and taxonomic synopsis of Cyperaceae in China. <em>Biodiversity Science.</em> 32(7): 24106., available online at https://doi.org/10.17520/biods.2024106 [details] Available for editors  PDF available [request]

 
 Present  Inaccurate  Introduced: alien  Containing type locality 
   

From editor or global species database
Additional information Chromosome number: 2n=70 [details]

Additional information Chromosome number: 2n=74 [details]

Additional information Chromosome number: 2n=82 [details]

Additional information Chromosome number: 2n=88 [details]

Description Rhizomes shortly creeping; shoots slender, markedly trigonous, 2–3 in each tuft; roots thick, pale yellow-brown; scales brown- or purple-red, usually persistent. Stems 30–120 cm, trigonous, but with a sharp, low ridge on the angles rough above, smooth below. Leaves up to 150 cm × 4–8 mm, serrulate at least towards tip, thin, rigid, plicate, gradually tapered to a fine point, mid green or yellow-green, quickly decaying; hypostomous; sheaths becoming purple-red, persistent, inner face hyaline, fibrillose on splitting, apex straight or concave; ligule 6–12 mm, acute. Inflorescence 1/4–1/3 length of stem; bracts leaf-like, lower exceeding inflorescence. Male spikes 2–4, 10–40 mm, the lower often with setaceous bracts; male glumes 4–6 mm, elliptic or oblanceolate, purplish-brown with green or paler midrib and hyaline margins; apex ± acute. Female spikes 2–3, ± contiguous but distant from the male spikes, 2–4(–7) cm, oblong-cylindric, erect, the lowest often pendent; female glumes 4–6 mm, narrowly lanceolate, purplish-brown with paler or green midrib; apex acute or acuminate, hyaline. Utricles 6–8 mm, ovoid-ellipsoid, inflated, ribbed, ± shiny, olive green to yellow-green, ascending; beak c. 2 mm, smooth, broadly notched; stigmas 3; nut obovoid, trigonous. [details]

Description Laxamente cespitosa, con rizoma robusto, de entrenudos algo largos. Tallos fértiles (20)35–100(110) cm, muy ásperos hacia el ápice, excepcionalmente lisos o casi cuadrangulares. Hojas (3)4–6(7) mm de anchura, de mayor longitud que los tallos, planas, muy ásperas en los bordes y nervio medio por el envés, ± rígidas; lígula (2)4–8 mm, de ápice obtuso; sin antelígula; vainas basales con limbo desarrollado, solo las más inferiores escuamiformes, de color pardo, raramente manchadas de color púrpura, enteras. Bráctea inferior folíacea, de mayor longitud que la inflorescencia, con frecuencia envainante. Espigas masculinas (1)2–3, de (8)20–40(55) mm, a veces andróginas, de lineares a estrechamente fusiformes; espigas femeninas 2–3, de (10)20–40(50) mm, generalmente cilíndricas, densifloras, la superior frecuentemente andrógina, distantes o las superiores aproximadas, pedunculadas, la inferior a veces colgante, el resto erectas. Glumas masculinas lineares a lanceoladas, agudas o subagudas, pardas, frecuentemente con margen escarioso en la zona apical; glumas femeninas lanceoladas a oblongas, obtusas, pardas o, excepcionalmente, negras, de longitud mucho mayor que los utrículos, de color pardo-rojizo, las de la zona media casi siempre con margen escarioso. Utrículos (3,9)4–6(8) × 1,2–3,2 mm, suberectos, ovoides o elipsoidales, inflados, pardo-amarillentos, brillantes, con los nervios prominentes, gradualmente atenuados en un pico de (1)1,1–2,2 mm, profundamente bífido, con el sinus ± agudo, liso. Aquenios 1,9–2,1 × 1–1,5 mm, de contorno oblongo o elíptico, trígonos, de color pardo. 2n = 74*, 82, 86*. [details]

Description Perennials. rhizome stoloniferous, stolons branched. Stems 30–100 cm tall, stout, triquetrous, scabrous on upper part. Leaves shorter than stem, 3–8 mm wide, pale green, scabrous. Basal sheaths reddish brown, ladder-like fibrillose. Involucral bracts leaflike, lowermost shortly sheathed, upper not sheathed. Inflorescences spikes 4–6; terminal 2 or 3 spikes male, approximate, linear cylindric, 2–3.5 cm long, subsessile; remaining spikes female, oblong or oblong-cylindric, 3–7 cm long, densely many flowered, shortly pedunculate. Staminate scales brown, apex obtuse to acute. Pistillate scales pale brown, midrib green, apex acute or acuminate. Perigynia yellow-green, obliquely patent, longer than pistillate scales, ovoid, inflated trigonous, 6–8 mm long, several veined, glabrous, long beaked, mouth bidentate. Achenes obovoid, trigonous, 1.7–2 mm long. Pistil style slender, flexuous; stigmas 3. [details]

Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 979 1753 to Sp. Pl. : 979 (1753), information provided by Alan E. on email Jun. 07 2021 More details could be found in The Plant L [details]

Translation Loosely cespitose, with a robust rhizome, with somewhat long internodes. Fertile stems (20)35–100(110) cm, very rough towards the apex, exceptionally smooth or almost quadrangular. Leaves (3)4–6(7) mm wide, longer than the stems, flat, very rough on the margins and on the midrib on the abaxial surface, ± rigid; ligule (2)4–8 mm, with an obtuse apex; anteligule absent; basal sheaths with a developed blade, only the lowest ones scale-like, brown in colour, rarely stained purple, entire. Lower bract leaf-like, longer than the inflorescence, frequently sheathing. Male spikes (1)2–3, (8)20–40(55) mm long, sometimes androgynous, linear to narrowly fusiform; female spikes 2–3, (10)20–40(50) mm long, generally cylindrical, densely flowered, the upper often androgynous, distant or the upper ones approximated, pedunculate, the lower sometimes drooping, the others erect. Male glumes linear to lanceolate, acute or subacute, brown, frequently with a scarious margin in the apical zone; female glumes lanceolate to oblong, obtuse, brown or, exceptionally, black, much longer than the utricles, reddish-brown, those of the middle zone almost always with a scarious margin. Utricles (3.9)4–6(8) × 1.2–3.2 mm, suberect, ovoid or ellipsoid, inflated, yellowish-brown, shiny, with prominent nerves, gradually attenuated into a beak (1)1.1–2.2 mm long, deeply bifid, with the sinus ± acute, smooth. Achenes 1.9–2.1 × 1–1.5 mm, oblong or elliptic in outline, trigonous, brown. 2n = 74*, 82, 86*. [details]
    Definitions

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LanguageName 
Japanese オニナルコスゲ(Oni-naruko-suge)  [details]