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Cyperaceae taxon details

Carex pseudocyperus L.

1678047  (urn:lsid:marinespecies.org:taxname:1678047)

accepted
Species

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  1. Variety Carex pseudocyperus var. haenkeana (C.Presl) Kük.
  2. Variety Carex pseudocyperus var. pseudocyperus
  3. Forma Carex pseudocyperus f. denticulata Y.L.Chang & Y.L.Yang (uncertain > unassessed)
  4. Subspecies Carex pseudocyperus subsp. acrifolia (V.I.Krecz.) Kukkonen (uncertain > unassessed)
  5. Subspecies Carex pseudocyperus subsp. afghanica Kukkonen (uncertain > unassessed)
  6. Variety Carex pseudocyperus var. solida T.J.Sørensen (uncertain > unassessed)
  7. Variety Carex pseudocyperus var. solida T.J.Sörensen (uncertain > unassessed)
  8. Forma Carex pseudocyperus f. minor Hampe accepted as Carex pseudocyperus var. pseudocyperus
  9. Forma Carex pseudocyperus f. multispicula Lepage accepted as Carex pseudocyperus var. pseudocyperus
  10. Forma Carex pseudocyperus f. novae-selandiae (Boeckeler) Kük. accepted as Carex fascicularis Sol. ex Boott
  11. Forma Carex pseudocyperus f. pallidior Kük. accepted as Carex pseudocyperus var. pseudocyperus
  12. Forma Carex pseudocyperus f. pseudocyperus L. accepted as Carex pseudocyperus L.
  13. Subspecies Carex pseudocyperus subsp. haenkeana (C.Presl) Kük. accepted as Carex pseudocyperus var. haenkeana (C.Presl) Kük.
  14. Subspecies Carex pseudocyperus subsp. platygluma C.B.Clarke ex Kük. accepted as Carex polystachya var. polystachya
  15. Variety Carex pseudocyperus var. americana Hochst. ex L.H.Bailey accepted as Carex pseudocyperus var. pseudocyperus
  16. Variety Carex pseudocyperus var. brachystachya Regel & Maack accepted as Carex capricornis Meinsh. ex Maxim.
  17. Variety Carex pseudocyperus var. cognata (Kunth) Boott accepted as Carex cognata Kunth
  18. Variety Carex pseudocyperus var. comosa (Boott) Boott accepted as Carex comosa Boott
  19. Variety Carex pseudocyperus var. fascicularis (Sol. ex Boott) Boott accepted as Carex fascicularis Sol. ex Boott
  20. Variety Carex pseudocyperus var. furcata Kük. accepted as Carex pseudocyperus var. pseudocyperus
  21. Variety Carex pseudocyperus var. interrupta L.C.Lamb. accepted as Carex pseudocyperus var. pseudocyperus
  22. Variety Carex pseudocyperus var. lechleri Boott accepted as Carex pseudocyperus var. haenkeana (C.Presl) Kük.
  23. Variety Carex pseudocyperus var. libanoticus Mouterde accepted as Carex pseudocyperus var. pseudocyperus
  24. Variety Carex pseudocyperus var. mazoviensis Zalewski accepted as Carex pseudocyperus var. pseudocyperus
  25. Variety Carex pseudocyperus var. polysticha (Boeckeler) Kük. accepted as Carex polysticha Boeckeler
terrestrial
Linnaeus, C. (1753). Species plantarum, exhibentes plantas rite cognitas ad genera relatas cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas. Vol 2. Salvius, Stockholm., available online at https://www.biodiversitylibrary.org/page/358580
page(s): 978 [details] 
Description Rhizomes short; shoots loosely tufted; roots thick (up to 1.5 mm), orange-brown, felted; scales dark brown or grey-brown,...  
Description Rhizomes short; shoots loosely tufted; roots thick (up to 1.5 mm), orange-brown, felted; scales dark brown or grey-brown, persistent. Stems 40–90 cm, solid, sharply trigonous; angles rough. Leaves up to 120 cm × 5–12 mm, rigid, erect, thin, plicate, very rough on keel and margins, tapered gradually to a fine point, bright yellow-green, passing through yellow to grey-brown on dying, persistent; hypostomous; sheaths becoming pink- or grey-brown, persistent, with inner face hyaline, forming fibrillae on splitting; apex straight or concave; ligule 10–15 mm, obtuse. Inflorescence 1/6–1/4 length of stem; bracts leaf-like, lowest 3–4 times longer than inflorescence. Male spike 1, 2–6 cm; male glumes 5–7 mm, elliptic-lanceolate, brown with a green or paler midrib; apex long-acuminate, ciliate. Female spikes 3–5, clustered at top and often exceeding male spike, 2–10 cm, cylindric, pendent; peduncles slender, rough, lowest only shortly ensheathed; female glumes 5–10 mm, ovate, brownish-hyaline, with midrib green with few fine hairs, drawn out into a long, fine, ciliate arista. Utricles 4–5 mm, broader than glumes, ovoid-ellipsoid, patent, ribbed, green, soon falling when ripe; beak c. 2 mm, smooth, deeply bifid; stigmas 3; nut obovoid, trigonous. [details]

Description Laxamente cespitosa, con rizoma muy robusto de entrenudos algo largos. Tallos fértiles (30)40–80(100) cm, escábridos en...  
Description Laxamente cespitosa, con rizoma muy robusto de entrenudos algo largos. Tallos fértiles (30)40–80(100) cm, escábridos en la zona superior, trígonos. Hojas (5)7–10(11) mm de anchura, de longitud mayor que los tallos, planas, muy ásperas en los bordes, algo rígidas; lígula 8–12(15) mm, de ápice obtuso, redondeado o subagudo; sin antelígula; vainas basales con limbo desarrollado, sólo las más inferiores escuamiformes, de color pardo claro, enteras. Bráctea inferior folíacea, de mayor longitud que la inflorescencia, envainante. Espiga masculina solitaria, (24)40–60(80) mm, lineal, a veces ginecandra o con algunos utrículos en el ápice y la base; espigas femeninas 3–5, de 20–50(70) mm, muy densas, aproximadas junto a la masculina a la que igualan o superan, colgantes, a veces la inferior algo separada del resto. Glumas masculinas estrechamente ovales, largamente mucronadas o aristadas, hialinas o pajizas, glumas femeninas lineares, largamente aristadas o mucronadas, de longitud igual o mayor —en ocasiones algo menor— que los utrículos, de color pardo-rojizo o pardo-púrpura oscuro, sin margen escarioso, con tres nervios bien marcados. Utrículos (3,7)4,5–5(6) × (0,9)1,2–1,5 mm, patentes o reflejos, de lineares a estrechamente ovales, ± trígonos, pajizos, con los nervios muy prominentes, gradualmente atenuados en un pico de 1,9–2,2 mm, profundamente bífido. Aquenios 1,4–1,8 × 0,8–1 mm, de contorno oval, trígonos, pajizos o de color pardo claro. 2n = 66*. [details]

Description Plants densely to loosely cespitose; rhizomes short, no more than 10 cm. Culms trigonous in cross section, 25–100 cm,...  
Description Plants densely to loosely cespitose; rhizomes short, no more than 10 cm. Culms trigonous in cross section, 25–100 cm, scabrous distally. Leaves: basal sheaths pale brown (rarely faintly red tinged); ligules usually much longer than wide; blades mid to dark green, flat to W-shaped, 4–13 mm wide, glabrous. Inflorescences 4–15 cm; proximal bract 12–55 cm, greatly exceeding inflorescence; proximal 2–5 spikes pistillate, erect or the proximal pendent, cylindric, 9–12 mm thick; terminal 1 spike staminate or, rarely, gynaecandrous, androgynous, or mixed. Pistillate scales lanceolate-acuminate, 2.5–8.6 × 0.3–0.6 mm, the distal shorter and the proximal longer than perigynia, margins ciliate, apex scabrous-awned. Staminate scales scabrous-awned, sometimes also ciliate-margined. Perigynia spreading to reflexed at maturity, strongly 12–20-veined, veins usually separated by less than 2 times their width, confluent at or proximal to mid beak (except for 2 prominent lateral), tightly investing achene, narrowly elliptic to lanceolate, 3.4–6.1 × 1–1.7 mm, leathery, apex gradually tapered; beak poorly defined, 1.2–2.2 mm, strongly bidentate, teeth straight or slightly out-curved, 0.7–1.2(–1.4) mm. Stigmas 3. Achenes pale brown, trigonous, smooth. 2n = 66.  [details]

Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 978 1753 to Sp. Pl. : 978 (1753), information provided by...  
Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 978 1753 to Sp. Pl. : 978 (1753), information provided by Alan E. on email Jun. 07 2021 More details could be found in The Plant L [details]
Cyperaceae Working Group. (2025). [see How to cite]. Global Cyperaceae Database. Carex pseudocyperus L.. Accessed at: https://cyperaceae.org/aphia.php?p=taxdetails&id=1678047 on 2026-05-30
Date
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2023-09-06 07:28:28Z
created
2024-12-10 11:47:54Z
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2025-03-11 11:19:03Z
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Nomenclature

original description Linnaeus, C. (1753). Species plantarum, exhibentes plantas rite cognitas ad genera relatas cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas. Vol 2. Salvius, Stockholm., available online at https://www.biodiversitylibrary.org/page/358580
page(s): 978 [details] 

basis of record Plants of the World Online (POWO). , available online at https://powo.science.kew.org/ [details] 

Taxonomy

taxonomy source Jermy, A.C.; Simpson, D.A.; Foley, M.J.Y.; Porter, M.S. (2007). Sedges of the British Isles. 554 pp. Botanical Society of Britain & Ireland, United Kingdom. [details] Available for editors  PDF available [request]

taxonomy source Luceño, M., Castroviejo, S., Jiménez-Mejías, P. (2008). Cyperaceae. In: Castroviejo, Luceño, M., Galán, A., Jiménez-Mejías, P., Cabezas, F., Medina, L. (eds). Flora iberica. Plantas vasculares de la Península Ibérica e Islas Baleares. Vol. XVIII. Madrid: CSIC., available online at https://bibdigital.rjb.csic.es/records/item/12958-flora-iberica-vol-18-cyperaceae-pontederiaceae?offset=, http://www.floraiberica.es/PHP/familias_lista_.php?familia=Cyperaceae [details] 

Other

additional source Li, B. Z.; Zhang, S. R. (2024). An updated species checklist and taxonomic synopsis of Cyperaceae in China. <em>Biodiversity Science.</em> 32(7): 24106., available online at https://doi.org/10.17520/biods.2024106 [details] Available for editors  PDF available [request]

 
 Present  Inaccurate  Introduced: alien  Containing type locality 
   

Syntype L 0052715 [details]
From editor or global species database
Additional information 2n=66 [details]

Description Rhizomes short; shoots loosely tufted; roots thick (up to 1.5 mm), orange-brown, felted; scales dark brown or grey-brown, persistent. Stems 40–90 cm, solid, sharply trigonous; angles rough. Leaves up to 120 cm × 5–12 mm, rigid, erect, thin, plicate, very rough on keel and margins, tapered gradually to a fine point, bright yellow-green, passing through yellow to grey-brown on dying, persistent; hypostomous; sheaths becoming pink- or grey-brown, persistent, with inner face hyaline, forming fibrillae on splitting; apex straight or concave; ligule 10–15 mm, obtuse. Inflorescence 1/6–1/4 length of stem; bracts leaf-like, lowest 3–4 times longer than inflorescence. Male spike 1, 2–6 cm; male glumes 5–7 mm, elliptic-lanceolate, brown with a green or paler midrib; apex long-acuminate, ciliate. Female spikes 3–5, clustered at top and often exceeding male spike, 2–10 cm, cylindric, pendent; peduncles slender, rough, lowest only shortly ensheathed; female glumes 5–10 mm, ovate, brownish-hyaline, with midrib green with few fine hairs, drawn out into a long, fine, ciliate arista. Utricles 4–5 mm, broader than glumes, ovoid-ellipsoid, patent, ribbed, green, soon falling when ripe; beak c. 2 mm, smooth, deeply bifid; stigmas 3; nut obovoid, trigonous. [details]

Description Laxamente cespitosa, con rizoma muy robusto de entrenudos algo largos. Tallos fértiles (30)40–80(100) cm, escábridos en la zona superior, trígonos. Hojas (5)7–10(11) mm de anchura, de longitud mayor que los tallos, planas, muy ásperas en los bordes, algo rígidas; lígula 8–12(15) mm, de ápice obtuso, redondeado o subagudo; sin antelígula; vainas basales con limbo desarrollado, sólo las más inferiores escuamiformes, de color pardo claro, enteras. Bráctea inferior folíacea, de mayor longitud que la inflorescencia, envainante. Espiga masculina solitaria, (24)40–60(80) mm, lineal, a veces ginecandra o con algunos utrículos en el ápice y la base; espigas femeninas 3–5, de 20–50(70) mm, muy densas, aproximadas junto a la masculina a la que igualan o superan, colgantes, a veces la inferior algo separada del resto. Glumas masculinas estrechamente ovales, largamente mucronadas o aristadas, hialinas o pajizas, glumas femeninas lineares, largamente aristadas o mucronadas, de longitud igual o mayor —en ocasiones algo menor— que los utrículos, de color pardo-rojizo o pardo-púrpura oscuro, sin margen escarioso, con tres nervios bien marcados. Utrículos (3,7)4,5–5(6) × (0,9)1,2–1,5 mm, patentes o reflejos, de lineares a estrechamente ovales, ± trígonos, pajizos, con los nervios muy prominentes, gradualmente atenuados en un pico de 1,9–2,2 mm, profundamente bífido. Aquenios 1,4–1,8 × 0,8–1 mm, de contorno oval, trígonos, pajizos o de color pardo claro. 2n = 66*. [details]

Description Plants densely to loosely cespitose; rhizomes short, no more than 10 cm. Culms trigonous in cross section, 25–100 cm, scabrous distally. Leaves: basal sheaths pale brown (rarely faintly red tinged); ligules usually much longer than wide; blades mid to dark green, flat to W-shaped, 4–13 mm wide, glabrous. Inflorescences 4–15 cm; proximal bract 12–55 cm, greatly exceeding inflorescence; proximal 2–5 spikes pistillate, erect or the proximal pendent, cylindric, 9–12 mm thick; terminal 1 spike staminate or, rarely, gynaecandrous, androgynous, or mixed. Pistillate scales lanceolate-acuminate, 2.5–8.6 × 0.3–0.6 mm, the distal shorter and the proximal longer than perigynia, margins ciliate, apex scabrous-awned. Staminate scales scabrous-awned, sometimes also ciliate-margined. Perigynia spreading to reflexed at maturity, strongly 12–20-veined, veins usually separated by less than 2 times their width, confluent at or proximal to mid beak (except for 2 prominent lateral), tightly investing achene, narrowly elliptic to lanceolate, 3.4–6.1 × 1–1.7 mm, leathery, apex gradually tapered; beak poorly defined, 1.2–2.2 mm, strongly bidentate, teeth straight or slightly out-curved, 0.7–1.2(–1.4) mm. Stigmas 3. Achenes pale brown, trigonous, smooth. 2n = 66.  [details]

Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 978 1753 to Sp. Pl. : 978 (1753), information provided by Alan E. on email Jun. 07 2021 More details could be found in The Plant L [details]

Translation Loosely tufted, with a very robust rhizome with somewhat long internodes. Fertile stems (30)40–80(100) cm tall, scabrid in the upper part, trigonous. Leaves (5)7–10(11) mm wide, longer than the stems, flat, very rough on the margins, somewhat rigid; ligule 8–12(15) mm, with an obtuse, rounded or subacute apex; without an antiligule; basal sheaths with a developed blade, only the lowermost scale-like, light brown in colour, entire. Lowest bract leaf-like, longer than the inflorescence, sheathing. Male spike solitary, (24)40–60(80) mm long, linear, sometimes gynecandrous or with a few utricles at the apex and base; female spikes 3–5, 20–50(70) mm long, very dense, closely grouped with the male spike, which they equal or exceed, pendulous, sometimes the lowest slightly separated from the rest. Male glumes narrowly oval, long-mucronate or aristate, hyaline or straw-coloured; female glumes linear, long-aristate or mucronate, as long as or longer—sometimes slightly shorter—than the utricles, reddish-brown to dark brown-purple in colour, without a scarious margin, with three well-marked nerves. Utricles (3.7)4.5–5(6) × (0.9)1.2–1.5 mm, spreading or reflexed, linear to narrowly oval, ± trigonous, straw-coloured, with very prominent nerves, gradually attenuated into a beak 1.9–2.2 mm long, deeply bifid. Achenes 1.4–1.8 × 0.8–1 mm, oval in outline, trigonous, straw-coloured or light brown. 2n = 66*. [details]
    Definitions

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LanguageName 
Japanese クグスゲ(Kugu-suge)  [details]