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Cyperaceae taxon details

Carex atrata subsp. atrata L.

To Field Museum Herbarium (Carex apodostachya Ohwi F1464064 isotype 1)
To Field Museum Herbarium (Carex apodostachya Ohwi F1464064 isotype 1)

1699757  (urn:lsid:marinespecies.org:taxname:1699757)

  1. Biota
  2. Plantae (Kingdom)
  3. Viridiplantae (Subkingdom)
  4. Streptophyta (Infrakingdom)
  5. Tracheophyta (Phylum (Division))
  6. Spermatophytina (Subphylum (Subdivision))
  7. Magnoliopsida (Class)
  8. Lilianae (Superorder)
  9. Poales (Order)
  10. Cyperaceae (Family)
  11. Cariceae (Tribe)
  12. Carex (Genus)
  13. Carex subg. Carex (Subgenus)
  14. Carex sect. Racemosae (Section)
  15. Carex atrata (Species)
  16. Carex atrata subsp. atrata (Subspecies)
accepted
Subspecies
Carex dubia Gaudin · unaccepted
terrestrial
Not documented
Description Laxamente cespitosa, con rizoma de entrenudos cortos. Tallos 13-26(35) cm, agudamente trígonos, lisos o algo escábridos,...  
Description Laxamente cespitosa, con rizoma de entrenudos cortos. Tallos 13-26(35) cm, agudamente trígonos, lisos o algo escábridos, densamente papilosos. Hojas (2)3-6(6,8) mm de anchura, de menor longitud que los tallos, ásperas en los bordes de la zona superior, planas, muy papilosas, glaucas, no muy rígidas; lígula hasta 2 mm, de ápice obtuso, redondeado o truncado; sin antelígula; vainas basales con limbo desarrollado; vainas basales escuamiformes, pardas o de color pardo oscuro o negruzcas. Bráctea inferior folíacea, de longitud mayor o menor que la inflorescencia, no cortamente envainante. Espigas (2)3-5(7), de 12-21 mm, pedunculadas, las inferiores colgantes o inclinadas, ovoides, ± aproximadas, la superior ginecandra, el resto femeninas o raramente todas ginecandras. Glumas ovales, agudas, negras; las masculinas frecuentemente con margen escarioso muy estrecho, pajizas, de mayor longitud que los utrículos. Estigmas 3. Utrículos 3-3,5 × 2-2,4 mm, suberectos, de contorno anchamente oval o suborbicular, comprimidos o plano-convexos, con los nervios no perceptibles, glabros, muy densamente papilosos, verdosos o de color pardo claro, variablemente teñidos de negro, bruscamente estrechados en un pico de 0,4-0,6 mm, recto, de ápice bidentado, papiloso. Aquenios 1,6-1,8 × 0,9-1 mm, de contorno elíptico u obovado, trígonos, verdosos o parduscos. 2n = 48-52*, 52*, 54*, 56*. [details]

Taxonomic remark Source in seed data: wcs Updated taxonomicStatus from Synonym to Accepted, information provided by Alan E. on email Jun. 03...  
Taxonomic remark Source in seed data: wcs Updated taxonomicStatus from Synonym to Accepted, information provided by Alan E. on email Jun. 03 2021 More details could be found in The Plant List v.1.1. Origin [details]
Cyperaceae Working Group. (2025). [see How to cite]. Global Cyperaceae Database. Carex atrata subsp. atrata L.. Accessed at: https://cyperaceae.org/aphia.php?p=taxdetails&id=1699757 on 2026-05-07
Date
action
by
2023-09-28 08:30:01Z
created
Jiménez-Mejías, Pedro
2024-12-10 11:47:54Z
unchecked
db_admin

Creative Commons License The webpage text is licensed under a Creative Commons Attribution 4.0 License


Nomenclature

basis of record Plants of the World Online (POWO). , available online at https://powo.science.kew.org/ [details] 

Taxonomy

taxonomy source Luceño, M., Castroviejo, S., Jiménez-Mejías, P. (2008). Cyperaceae. In: Castroviejo, Luceño, M., Galán, A., Jiménez-Mejías, P., Cabezas, F., Medina, L. (eds). Flora iberica. Plantas vasculares de la Península Ibérica e Islas Baleares. Vol. XVIII. Madrid: CSIC., available online at https://bibdigital.rjb.csic.es/records/item/12958-flora-iberica-vol-18-cyperaceae-pontederiaceae?offset=, http://www.floraiberica.es/PHP/familias_lista_.php?familia=Cyperaceae [details] 

Other

additional source Li, B. Z.; Zhang, S. R. (2024). An updated species checklist and taxonomic synopsis of Cyperaceae in China. <em>Biodiversity Science.</em> 32(7): 24106., available online at https://doi.org/10.17520/biods.2024106 [details] Available for editors  PDF available [request]

 
 Present  Inaccurate  Introduced: alien  Containing type locality 
   

From editor or global species database
Description Laxamente cespitosa, con rizoma de entrenudos cortos. Tallos 13-26(35) cm, agudamente trígonos, lisos o algo escábridos, densamente papilosos. Hojas (2)3-6(6,8) mm de anchura, de menor longitud que los tallos, ásperas en los bordes de la zona superior, planas, muy papilosas, glaucas, no muy rígidas; lígula hasta 2 mm, de ápice obtuso, redondeado o truncado; sin antelígula; vainas basales con limbo desarrollado; vainas basales escuamiformes, pardas o de color pardo oscuro o negruzcas. Bráctea inferior folíacea, de longitud mayor o menor que la inflorescencia, no cortamente envainante. Espigas (2)3-5(7), de 12-21 mm, pedunculadas, las inferiores colgantes o inclinadas, ovoides, ± aproximadas, la superior ginecandra, el resto femeninas o raramente todas ginecandras. Glumas ovales, agudas, negras; las masculinas frecuentemente con margen escarioso muy estrecho, pajizas, de mayor longitud que los utrículos. Estigmas 3. Utrículos 3-3,5 × 2-2,4 mm, suberectos, de contorno anchamente oval o suborbicular, comprimidos o plano-convexos, con los nervios no perceptibles, glabros, muy densamente papilosos, verdosos o de color pardo claro, variablemente teñidos de negro, bruscamente estrechados en un pico de 0,4-0,6 mm, recto, de ápice bidentado, papiloso. Aquenios 1,6-1,8 × 0,9-1 mm, de contorno elíptico u obovado, trígonos, verdosos o parduscos. 2n = 48-52*, 52*, 54*, 56*. [details]

Taxonomic remark Source in seed data: wcs Updated taxonomicStatus from Synonym to Accepted, information provided by Alan E. on email Jun. 03 2021 More details could be found in The Plant List v.1.1. Origin [details]

Translation Loosely cespitose, with short-internode rhizomes. Stems 13–26(35) cm, sharply trigonous, smooth or slightly scabrid, densely papillose. Leaves (2)3–6(6.8) mm wide, shorter than the stems, rough along the margins of the upper part, flat, very papillose, glaucous, not very rigid; ligule up to 2 mm, with obtuse, rounded or truncate apex; no antiligule; basal sheaths with developed blade; basal scale-like sheaths brown, dark brown or blackish. Lower bract leaf-like, shorter or longer than the inflorescence, not shortly sheathing. Spikes (2)3–5(7), 12–21 mm long, pedunculate, the lower ones pendent or inclined, ovoid, ± approximate; the upper spike gynandrous, the remainder female or rarely all gynandrous. Glumes oval, acute, black; male glumes frequently with a very narrow scarious margin, straw-coloured, longer than the utricles. Stigmas 3. Utricles 3–3.5 × 2–2.4 mm, suberect, broadly oval to suborbicular in outline, compressed or plano-convex, with nerves not visible, glabrous, very densely papillose, greenish or light brown, variably black-tinged, abruptly narrowed into a straight beak 0.4–0.6 mm long, with a bidentate, papillose apex. Achenes 1.6–1.8 × 0.9–1 mm, elliptic or obovate in outline, trigonous, greenish or brownish. 2n = 48–52*, 52*, 54*, 56*. [details]
    Definitions

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To Biodiversity Heritage Library (1 publication) (from synonym Carex aethostachya Schkuhr)
To Biodiversity Heritage Library (1 publication) (from synonym Carex japonalpina (T.Koyama) T.Koyama)
To Biodiversity Heritage Library (1 publication) (from synonym Carex pseudocilicica Hausskn.)
To Biodiversity Heritage Library (1 publication) (from synonym Carex atrata var. bicolor Čelak.)
To Biodiversity Heritage Library (109 publications) (from synonym Carex castanea Miel. ex Saut.)
To Biodiversity Heritage Library (120 publications) (from synonym Carex atrofusca Steven)
To Biodiversity Heritage Library (20 publications) (from synonym Carex atropurpurea Fisch. ex Trevir.)
To Biodiversity Heritage Library (21 publications) (from synonym Carex dubia Gaudin)
To Biodiversity Heritage Library (3 publications) (from synonym Carex paishanensis Nakai)
To Biodiversity Heritage Library (3 publications) (from synonym Carex atrata var. rectiuscula Hartm.)
To Biodiversity Heritage Library (44 publications) (from synonym Carex cylindrica Miel. ex Kunth)
To European Nucleotide Archive, ENA (Carex atrofusca) (from synonym Carex atrofusca Steven)
To European Nucleotide Archive, ENA (Carex castanea) (from synonym Carex castanea Miel. ex Saut.)
To European Nucleotide Archive, ENA (Carex japonalpina) (from synonym Carex japonalpina (T.Koyama) T.Koyama)
To Field Museum Herbarium (Carex apodostachya Ohwi F1464064 isotype 1) (from synonym Carex apodostachya Ohwi)
To GenBank (3 nucleotides; 1 proteins) (from synonym Carex japonalpina (T.Koyama) T.Koyama)
To International Plant Names Index (IPNI) (from synonym Carex japonalpina (T.Koyama) T.Koyama)
To International Plant Names Index (IPNI) (from synonym Carex atropurpurea Fisch. ex Trevir.)
To International Plant Names Index (IPNI) (from synonym Carex cylindrica Miel. ex Kunth)
To International Plant Names Index (IPNI) (from synonym Carex apodostachya Ohwi)
To International Plant Names Index (IPNI) (from synonym Carex aethostachya Schkuhr)
To International Plant Names Index (IPNI) (from synonym Carex paishanensis Nakai)
To International Plant Names Index (IPNI) (from synonym Carex tauricola Boiss.)
To International Plant Names Index (IPNI) (from synonym Carex dubia Gaudin)
To International Plant Names Index (IPNI) (from synonym Carex castanea Miel. ex Saut.)
To International Plant Names Index (IPNI) (from synonym Carex atrofusca Steven)
To International Plant Names Index (IPNI) 
To International Plant Names Index (IPNI) (from synonym Carex bigelowii subsp. paishanensis (Nakai) Vorosch.)
To International Plant Names Index (IPNI) (from synonym Carex atrata subsp. apodostachya (Ohwi) T.Koyama)
To International Plant Names Index (IPNI) (from synonym Carex atrata subsp. castanea (Nyman) K.Richt.)
To International Plant Names Index (IPNI) (from synonym Carex nigra subsp. distachya (Nyman) K.Richt.)
To International Plant Names Index (IPNI) (from synonym Carex atrata var. brunnescens Andersson)
To International Plant Names Index (IPNI) (from synonym Carex atrata var. japonoalpina T.Koyama)
To International Plant Names Index (IPNI) (from synonym Carex atrata var. ovata Boott)
To International Plant Names Index (IPNI) (from synonym Carex perfusca var. japonoalpina (T.Koyama) Kitag.)
To International Plant Names Index (IPNI) (from synonym Carex atrata var. heterostachya Almq.)
To International Plant Names Index (IPNI) (from synonym Carex pseudocilicica Hausskn.)
To International Plant Names Index (IPNI) (from synonym Carex atrata var. bicolor Čelak.)
To International Plant Names Index (IPNI) (from synonym Carex atrata var. rectiuscula Hartm.)
To International Plant Names Index (IPNI) (from synonym Carex nigra var. distachya Nyman)
To International Plant Names Index (IPNI) (from synonym Carex atrata var. pseudocilicica Kük.)
To International Plant Names Index (IPNI) (from synonym Carex atrata var. spiculosior Norman)
To International Plant Names Index (IPNI) (from synonym Carex atrata var. spadicea Beurl.)
To International Plant Names Index (IPNI) (from synonym Carex atrata var. dubia Gaudin)
To International Plant Names Index (IPNI) (from synonym Carex atrata var. laxa Neilr.)
To International Plant Names Index (IPNI) (from synonym Carex atrata var. castanea Nyman)
To Photos of Cyperaceae by Hoshino Labo. from Japan (from synonym Carex atrata var. japonoalpina T.Koyama)
To Plants of the World Online (from synonym Carex atrata var. laxa Neilr.)
To Plants of the World Online (from synonym Carex atrata var. japonoalpina T.Koyama)
To Plants of the World Online (from synonym Carex nigra var. distachya Nyman)
To Plants of the World Online (from synonym Carex atrata var. castanea Nyman)
To Plants of the World Online (from synonym Carex atrata var. spiculosior Norman)
To Plants of the World Online (from synonym Carex atrata f. castanea (Nyman) K.Richt.)
To Plants of the World Online (from synonym Carex atrata f. decolorans Neuman)
To Plants of the World Online (from synonym Carex atrata f. gelida Schur)
To Plants of the World Online (from synonym Carex atrata var. rectiuscula Hartm.)
To Plants of the World Online (from synonym Carex atrata var. pseudocilicica Kük.)
To Plants of the World Online (from synonym Carex atrata var. spadicea Beurl.)
To Plants of the World Online (from synonym Carex atrata var. bicolor Čelak.)
To Plants of the World Online (from synonym Carex atrata var. heterostachya Almq.)
To Plants of the World Online (from synonym Carex perfusca var. japonoalpina (T.Koyama) Kitag.)
To Plants of the World Online (from synonym Carex atrata var. dubia Gaudin)
To Plants of the World Online (from synonym Carex atrata var. brunnescens Andersson)
To Plants of the World Online (from synonym Carex atrata subsp. bicolor Čelak.)
To Plants of the World Online (from synonym Carex atrata var. ovata Boott)
To Plants of the World Online (from synonym Carex nigra subsp. distachya (Nyman) K.Richt.)
To Plants of the World Online (from synonym Carex atrata subsp. castanea (Nyman) K.Richt.)
To Plants of the World Online (from synonym Carex atrata subsp. apodostachya (Ohwi) T.Koyama)
To Plants of the World Online (from synonym Carex atrata subforma aristata Podp.)
To Plants of the World Online (from synonym Carex bigelowii subsp. paishanensis (Nakai) Vorosch.)
To Plants of the World Online 
To Plants of the World Online (from synonym Carex pseudocilicica Hausskn.)
To Plants of the World Online (from synonym Carex tauricola Boiss.)
To Plants of the World Online (from synonym Carex dubia Gaudin)
To Plants of the World Online (from synonym Carex atrofusca Steven)
To Plants of the World Online (from synonym Carex castanea Miel. ex Saut.)
To Plants of the World Online (from synonym Carex paishanensis Nakai)
To Plants of the World Online (from synonym Carex aethostachya Schkuhr)
To Plants of the World Online (from synonym Carex apodostachya Ohwi)
To Plants of the World Online (from synonym Carex atropurpurea Fisch. ex Trevir.)
To Plants of the World Online (from synonym Carex cylindrica Miel. ex Kunth)
To Plants of the World Online (from synonym Carex japonalpina (T.Koyama) T.Koyama)
To Royal Botanic Gardens, Kew (Carex apodostachya K000961019 type 1) (from synonym Carex apodostachya Ohwi)
To University of Tokyo Herbarium (Carex  atrata subsp.  perfusca  var.  japonalpina type TI00010005) (from synonym Carex atrata var. japonoalpina T.Koyama)
To University of Tokyo Herbarium (Carex  atrata subsp.  perfusca  var.  japonalpina type TI00010006) (from synonym Carex atrata var. japonoalpina T.Koyama)
To University of Tokyo Herbarium (Carex  atrata subsp.  perfusca  var.  japonalpina type TI00010007) (from synonym Carex atrata var. japonoalpina T.Koyama)
To University of Tokyo Herbarium (Carex  atrata subsp.  perfusca  var.  japonalpina type TI00010008) (from synonym Carex atrata var. japonoalpina T.Koyama)
To University of Tokyo Herbarium (Carex  atrata subsp.  perfusca  var.  japonalpina type TI00010009) (from synonym Carex atrata var. japonoalpina T.Koyama)
To University of Tokyo Herbarium (Carex  atrata subsp.  perfusca  var.  japonalpina type TI00010011) (from synonym Carex atrata var. japonoalpina T.Koyama)
To University of Tokyo Herbarium (Carex  atrata subsp.  perfusca  var.  japonalpina type TI00010012) (from synonym Carex atrata var. japonoalpina T.Koyama)
To University of Tokyo Herbarium (Carex  atrata subsp.  perfusca  var.  japonalpina type TI00010013) (from synonym Carex atrata var. japonoalpina T.Koyama)
To University of Tokyo Herbarium (Carex  atrata subsp.  perfusca  var.  japonalpina type TI00010014) (from synonym Carex atrata var. japonoalpina T.Koyama)
To University of Tokyo Herbarium (Carex  atrata subsp.  perfusca  var.  japonalpina type TI00010015) (from synonym Carex atrata var. japonoalpina T.Koyama)
To University of Tokyo Herbarium (Carex  atrata subsp.  perfusca  var.  japonalpina type TI00010016) (from synonym Carex atrata var. japonoalpina T.Koyama)
To University of Tokyo Herbarium (Carex  atrata subsp.  perfusca  var.  japonalpina type TI00010100) (from synonym Carex atrata var. japonoalpina T.Koyama)
To University of Tokyo Herbarium (Carex  paishanensis  type TI00010189) (from synonym Carex paishanensis Nakai)
From editor or global species database
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