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Cyperaceae taxon details

Carex arenaria L.

438828  (urn:lsid:marinespecies.org:taxname:438828)

  1. Biota
  2. Plantae (Kingdom)
  3. Viridiplantae (Subkingdom)
  4. Streptophyta (Infrakingdom)
  5. Tracheophyta (Phylum (Division))
  6. Spermatophytina (Subphylum (Subdivision))
  7. Magnoliopsida (Class)
  8. Lilianae (Superorder)
  9. Poales (Order)
  10. Cyperaceae (Family)
  11. Cariceae (Tribe)
  12. Carex (Genus)
  13. Carex subg. Vignea (Subgenus)
  14. Carex sect. Ammoglochin (Section)
  15. Carex arenaria (Species)
accepted
Species

Ordering

  • Alphabetically
  • By status

Children Display

  1. Forma Carex arenaria f. pallida Lackow. accepted as Carex arenaria L.
  2. Forma Carex arenaria f. pumila Lackow. accepted as Carex arenaria L.
  3. Forma Carex arenaria f. remota (T.Marsson) Almq. accepted as Carex arenaria L.
  4. Subspecies Carex arenaria subsp. colchica (J.Gay) Nyman accepted as Carex colchica J.Gay
  5. Subspecies Carex arenaria subsp. ligerica (J.Gay) Bonnier & Layens accepted as Carex colchica J.Gay
  6. Subspecies Carex arenaria subsp. repens (Bellardi) Wahlenb. accepted as Carex repens Bellardi
  7. Variety Carex arenaria var. adjunta Merino accepted as Carex arenaria L.
  8. Variety Carex arenaria var. castanea Boott accepted as Carex colchica J.Gay
  9. Variety Carex arenaria var. colchica (J.Gay) Christ accepted as Carex colchica J.Gay
  10. Variety Carex arenaria var. posnaniensis (Sprib.) Asch. & Graebn. accepted as Carex repens Bellardi
  11. Variety Carex arenaria var. prostrata De Langhe accepted as Carex arenaria L.
  12. Variety Carex arenaria var. pseudoarenaria T.Marsson accepted as Carex arenaria L.
  13. Variety Carex arenaria var. pseudobrizoides (Clavaud) Nyman accepted as Carex pseudobrizoides Clavaud
  14. Variety Carex arenaria var. remota T.Marsson accepted as Carex arenaria L.
  15. Variety Carex arenaria var. repens (Bellardi) Wahlenb. accepted as Carex repens Bellardi
  16. Variety Carex arenaria var. spiralis Asch. & Graebn. accepted as Carex arenaria L.
  17. Variety Carex arenaria var. tenuis Asch. & Graebn. accepted as Carex arenaria L.
  18. Variety Carex arenaria var. umbrosa Coss. & Germ. accepted as Carex pseudobrizoides Clavaud
  19. Variety Carex arenaria var. umbrosa (Sprib.) Sprib. accepted as Carex repens Bellardi
marine, terrestrial
Not documented
Description Rhizomes far-creeping; shoots usually single, at about every fourth node; roots pale brown, much branched; scales dark brown,...  
Description Rhizomes far-creeping; shoots usually single, at about every fourth node; roots pale brown, much branched; scales dark brown, soon becoming fibrous. Stems 10–90 cm, rough towards the top, trigonous, often curved, varying in thickness. Leaves up to 60 cm × 1.5–3.5 mm, ± flat, rigid, thick, rough above, often recurved and keeled or channelled in open habitats, tapering gradually to a fine trigonous point, shiny, dark green, often dark brown on dying; base frequently with a small scarious flange on either side; hypostomous; lower sheaths pale brown or grey-brown, persistent, with inner face a loose hyaline membrane, densely mamillate (appearing as a velvet sheen), becoming brown; apex straight; ligule 3–5 mm, obtuse, tubular. Inflorescence a dense head up to 8 cm; bracts glumaceous, lower with setaceous points. Spikes 5–15, 8–14 mm, terminal male, middle male at top and female below, lower entirely female. Male glumes 5–7 mm, lanceolate-elliptic, pale red-brown with hyaline margins; apex acute. Female glumes 5–6 mm, ovate, pale red-brown, with pale or green midrib and hyaline margins; apex acute or acuminate. Utricles 4–5.5 mm, ovate, plano-convex, many-ribbed, broadly winged, serrate in upper half, pale green-brown; beak 1–1.5 mm, broadly winged below, bifid; stigmas 2; nut oblong-ellipsoid. [details]

Description Rizoma de entrenudos muy largos, total o parcialmente cubierto por escamas que surgen de los nudos, de color pardo,...  
Description Rizoma de entrenudos muy largos, total o parcialmente cubierto por escamas que surgen de los nudos, de color pardo, deshechas en fibras en su parte proximal, raramente enteras. Tallos (8)15–30(40) cm, trígonos, con los ángulos agudos, escábridos en la parte superior. Hojas (1)2–3,5(5) mm de anchura, de menor longitud que los tallos, planas, canaliculadas, plegadas o enrolladas, en este último caso pudiendo ser estrechas (0,4 mm), ásperas en los bordes, no muy rígidas; lígula (0,8)1–2(3) mm, con los bordes levantados, de anchura menor que el limbo, de ápice obtuso o redondeado y borde generalmente ciliado; sin antelígula; vainas basales subterráneas, iguales a las escamas del rizoma, las aéreas generalmente con limbos desarrollados. Bráctea inferior de longitud mucho menor que la inflorescencia, setácea, rara vez cortamente foliácea, con vaina abierta en su base. Espigas dispuestas en espiga simple (1,8)4–6(7) cm, agrupadas, la superior frecuentemente masculina, las medias andróginas y la inferior generalmente femenina. Glumas ovales, acuminadas o aristadas, de color pardo claro cuando jóvenes y pardo rojizo ± oscuro en la madurez, las femeninas mayores que los utrículos. Utrículos (4)4,5–5,5(8) × (1,1)1,5–2(2) mm, de contorno lanceolado u oval, plano-convexos o cóncavo-convexos, esponjosos en la base, con la ancha maya presente a lo largo de la mayor parte de la longitud de cada uno de sus bordes, prominentemente nerviados, oliváceos o de color pardo obscuro en la madurez, gradualmente atenuados en un pico de 1,2–1,7(2) mm, escábrido-serrulado, bífido. Aquenios (1,4)1,7–2,1 × (0,6)0,9–1,1(1,2) mm, de contorno oblongo, obovado o elíptico, cóncavos menos que convexos, con la base del estilo persistente en forma de una corta columna. 2n = 58, 60*, 64*. [details]

Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 2: 973. 1753 [1 May 1753] to Sp. Pl. : 973 (1753),...  
Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 2: 973. 1753 [1 May 1753] to Sp. Pl. : 973 (1753), information provided by Alan E. on email Jun. 07 2021 More details could be found in [details]
Cyperaceae Working Group. (2025). [see How to cite]. Global Cyperaceae Database. Carex arenaria L.. Accessed at: https://www.cyperaceae.org/aphia.php?p=taxdetails&id=438828 on 2026-05-05
Date
action
by
2009-12-10 15:07:16Z
created
Decock, Wim
2023-09-05 10:00:04Z
changed
Jiménez-Mejías, Pedro
2024-12-10 11:47:54Z
unchecked
db_admin
2025-03-11 11:19:03Z
changed
Márquez-Corro, José Ignacio

Creative Commons License The webpage text is licensed under a Creative Commons Attribution 4.0 License


Nomenclature

basis of record Poma, L. (1936). Botanisch verslag van de excursies naar het Zwin op 15 mei en 13 september 1936. Biol. Jb. Dodonaea 3: 65-69 [details] 

Taxonomy

taxonomy source Jermy, A.C.; Simpson, D.A.; Foley, M.J.Y.; Porter, M.S. (2007). Sedges of the British Isles. 554 pp. Botanical Society of Britain & Ireland, United Kingdom. [details] Available for editors  PDF available [request]

taxonomy source Luceño, M., Castroviejo, S., Jiménez-Mejías, P. (2008). Cyperaceae. In: Castroviejo, Luceño, M., Galán, A., Jiménez-Mejías, P., Cabezas, F., Medina, L. (eds). Flora iberica. Plantas vasculares de la Península Ibérica e Islas Baleares. Vol. XVIII. Madrid: CSIC., available online at https://bibdigital.rjb.csic.es/records/item/12958-flora-iberica-vol-18-cyperaceae-pontederiaceae?offset=, http://www.floraiberica.es/PHP/familias_lista_.php?familia=Cyperaceae [details] 

Other

context source (RAS) Australian Antarctic Data Centre. , available online at https://data.aad.gov.au/aadc/biodiversity/ [details] 

additional source Plants of the World Online (POWO). , available online at https://powo.science.kew.org/ [details] 

 
 Present  Inaccurate  Introduced: alien  Containing type locality 
   

From editor or global species database
Description Rhizomes far-creeping; shoots usually single, at about every fourth node; roots pale brown, much branched; scales dark brown, soon becoming fibrous. Stems 10–90 cm, rough towards the top, trigonous, often curved, varying in thickness. Leaves up to 60 cm × 1.5–3.5 mm, ± flat, rigid, thick, rough above, often recurved and keeled or channelled in open habitats, tapering gradually to a fine trigonous point, shiny, dark green, often dark brown on dying; base frequently with a small scarious flange on either side; hypostomous; lower sheaths pale brown or grey-brown, persistent, with inner face a loose hyaline membrane, densely mamillate (appearing as a velvet sheen), becoming brown; apex straight; ligule 3–5 mm, obtuse, tubular. Inflorescence a dense head up to 8 cm; bracts glumaceous, lower with setaceous points. Spikes 5–15, 8–14 mm, terminal male, middle male at top and female below, lower entirely female. Male glumes 5–7 mm, lanceolate-elliptic, pale red-brown with hyaline margins; apex acute. Female glumes 5–6 mm, ovate, pale red-brown, with pale or green midrib and hyaline margins; apex acute or acuminate. Utricles 4–5.5 mm, ovate, plano-convex, many-ribbed, broadly winged, serrate in upper half, pale green-brown; beak 1–1.5 mm, broadly winged below, bifid; stigmas 2; nut oblong-ellipsoid. [details]

Description Rizoma de entrenudos muy largos, total o parcialmente cubierto por escamas que surgen de los nudos, de color pardo, deshechas en fibras en su parte proximal, raramente enteras. Tallos (8)15–30(40) cm, trígonos, con los ángulos agudos, escábridos en la parte superior. Hojas (1)2–3,5(5) mm de anchura, de menor longitud que los tallos, planas, canaliculadas, plegadas o enrolladas, en este último caso pudiendo ser estrechas (0,4 mm), ásperas en los bordes, no muy rígidas; lígula (0,8)1–2(3) mm, con los bordes levantados, de anchura menor que el limbo, de ápice obtuso o redondeado y borde generalmente ciliado; sin antelígula; vainas basales subterráneas, iguales a las escamas del rizoma, las aéreas generalmente con limbos desarrollados. Bráctea inferior de longitud mucho menor que la inflorescencia, setácea, rara vez cortamente foliácea, con vaina abierta en su base. Espigas dispuestas en espiga simple (1,8)4–6(7) cm, agrupadas, la superior frecuentemente masculina, las medias andróginas y la inferior generalmente femenina. Glumas ovales, acuminadas o aristadas, de color pardo claro cuando jóvenes y pardo rojizo ± oscuro en la madurez, las femeninas mayores que los utrículos. Utrículos (4)4,5–5,5(8) × (1,1)1,5–2(2) mm, de contorno lanceolado u oval, plano-convexos o cóncavo-convexos, esponjosos en la base, con la ancha maya presente a lo largo de la mayor parte de la longitud de cada uno de sus bordes, prominentemente nerviados, oliváceos o de color pardo obscuro en la madurez, gradualmente atenuados en un pico de 1,2–1,7(2) mm, escábrido-serrulado, bífido. Aquenios (1,4)1,7–2,1 × (0,6)0,9–1,1(1,2) mm, de contorno oblongo, obovado o elíptico, cóncavos menos que convexos, con la base del estilo persistente en forma de una corta columna. 2n = 58, 60*, 64*. [details]

Taxonomic remark Source in seed data: wcs Update namepublishedIn from Sp. Pl. 2: 973. 1753 [1 May 1753] to Sp. Pl. : 973 (1753), information provided by Alan E. on email Jun. 07 2021 More details could be found in [details]

Translation Rhizome with very long internodes, totally or partially covered by brown scales arising from the nodes, disintegrating into fibres in their proximal part, rarely entire. Stems (8)15–30(40) cm, trigonous, with sharp angles, scabrid in the upper part. Leaves (1)2–3.5(5) mm wide, shorter than the stems, flat, channelled, folded or rolled, in the latter case sometimes narrow (0.4 mm), rough on the margins, not very rigid; ligule (0.8)1–2(3) mm, with raised margins, narrower than the blade, with an obtuse or rounded apex and a generally ciliate margin; without an anteligule; basal sheaths subterranean, similar to the rhizome scales, the aerial ones generally with well-developed blades. Lower bract much shorter than the inflorescence, setaceous, rarely shortly foliaceous, with an open sheath at the base. Spikes arranged in a simple spike (1.8)4–6(7) cm, clustered, the upper one often male, the middle ones androgynous and the lower one usually female. Glumes ovate, acuminate or aristate, light brown when young and reddish-brown ± dark at maturity, the female ones longer than the utricles. Utricles (4)4.5–5.5(8) × (1.1)1.5–2(2) mm, lanceolate or ovate in outline, plano-convex or concavo-convex, spongy at the base, with a broad wing present along most of the length of each margin, prominently nerved, olive-coloured or dark brown at maturity, gradually attenuate into a beak 1.2–1.7(2) mm long, scabrid-serrulate, bifid. Achenes (1.4)1.7–2.1 × (0.6)0.9–1.1(1.2) mm, oblong, obovoid or elliptic in outline, less concave than convex, with the style base persistent as a short column. 2n = 58, 60*, 64*. [details]
    Definitions

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LanguageName 
Dutch zandzegge  [details]
English sand sedge  [details]
Norwegian Bokmål sandstarr  [details]
Norwegian Nynorsk sandstorr  [details]
Scottish Gaelic cíb ghainimh  [details]
Swedish sandstarr  [details]
Welsh hesgen y tywod  [details]
To Barcode of Life (33 barcodes) 
To Biodiversity Heritage Library (1 publication) (from synonym Carex arenaria var. tenuis Asch. & Graebn.)
To Biodiversity Heritage Library (1449 publications) 
To Biodiversity Heritage Library (18 publications) (from synonym Carex schreberi Desv.)
To Biodiversity Heritage Library (28 publications) (from synonym Vignea arenaria (L.) Rchb.)
To Biodiversity Heritage Library (298 publications) (from synonym Carex intermedia d'Urv.)
To Biodiversity Heritage Library (75 publications) (from synonym Carex spadicea Gilib.)
To European Nucleotide Archive, ENA (Carex arenaria) 
To International Plant Names Index (IPNI) 
To International Plant Names Index (IPNI) (from synonym Vignea arenaria (L.) Rchb.)
To International Plant Names Index (IPNI) (from synonym Carex intermedia d'Urv.)
To International Plant Names Index (IPNI) (from synonym Carex sabuletorum M.Bieb. ex Boott)
To International Plant Names Index (IPNI) (from synonym Carex schreberi Desv.)
To International Plant Names Index (IPNI) (from synonym Carex spadicea Gilib.)
To International Plant Names Index (IPNI) (from synonym Carex witheringii Gray)
To International Plant Names Index (IPNI) (from synonym Carex arenaria var. prostrata De Langhe)
To International Plant Names Index (IPNI) (from synonym Carex arenaria var. spiralis Asch. & Graebn.)
To International Plant Names Index (IPNI) (from synonym Carex arenaria var. pseudoarenaria T.Marsson)
To International Plant Names Index (IPNI) (from synonym Carex arenaria var. remota T.Marsson)
To International Plant Names Index (IPNI) (from synonym Carex arenaria var. adjunta Merino)
To International Plant Names Index (IPNI) (from synonym Carex arenaria var. tenuis Asch. & Graebn.)
To NHMUK collection (Carex arenaria L.; NHMUK:ecatalogue:9094262) 
To Plants of the World Online 
To Plants of the World Online (from synonym Carex intermedia d'Urv.)
To Plants of the World Online (from synonym Vignea arenaria (L.) Rchb.)
To Plants of the World Online (from synonym Carex schreberi Desv.)
To Plants of the World Online (from synonym Carex spadicea Gilib.)
To Plants of the World Online (from synonym Carex sabuletorum M.Bieb. ex Boott)
To Plants of the World Online (from synonym Carex arenaria var. spiralis Asch. & Graebn.)
To Plants of the World Online (from synonym Carex arenaria var. remota T.Marsson)
To Plants of the World Online (from synonym Carex arenaria var. tenuis Asch. & Graebn.)
To Plants of the World Online (from synonym Carex arenaria var. adjunta Merino)
To Plants of the World Online (from synonym Carex arenaria var. pseudoarenaria T.Marsson)
To Plants of the World Online (from synonym Carex arenaria var. prostrata De Langhe)
To Plants of the World Online (from synonym Carex witheringii Gray)
To Plants of the World Online (from synonym Carex arenaria f. pallida Lackow.)
To Plants of the World Online (from synonym Carex arenaria f. pumila Lackow.)
To Plants of the World Online (from synonym Carex arenaria f. remota (T.Marsson) Almq.)
Unreviewed
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